




REVISIONARY CLASSIFICATION OF RUTILIINI 69 
male genitalia are examined (after removal from the fly), and there appear to be no 
intermediates. It has therefore been concluded that ‘imperialis’ of authors consists 
of a confusion of two very closely allied species which are indistinguishable on 
external characters (or apparently so as none have been found of any value). As 
Guérin-Méneville’s type-material (which included a male specimen) is lost it was 
uncertain to which of the two species his name imperialis should be applied; this has 
therefore been resolved by neotype designation (see p. 125). The species with the 
prominence on the surstylus and the very slender cerci appears to be more common 
than the other species and the neotype specimen chosen for imperialis (which shows 
the genital characters clearly although the hypopygium remains in situ) is therefore 
of this species; there is no available name for the second species, which is therefore 
here described as new with the name imperialoides. The new species is clearly 
extremely closely allied to imperialis and examination of male genitalia is essential 
for accurate identification of imperialoides and imperialis. 
Subgenus DONOVANIUS Enderlein stat. n. 
Donovanius Enderlein, 1936 : 409. Type-species: Musca regalis Guérin-Méneville, 1831, by 
original designation. 
Psavonia Enderlein, 1936 : 414. Type-species: Psaronia bisetosa Enderlein, 1936, by original 
designation. Syn. n. 
Menevillea Enderlein, 1936 : 416. Type-species: Rutilia pellucens Macquart, 1846, by original 
designation [presumed correctly identified, but see discussion]. Syn. n. 
Diacnosis. Parafrontals pollinose, not metallic. Epistome and genae non-metallic. Facial 
carina flattened, slightly widened at level of base of third antennal segment, merging into lunula 
without a distinct depression. Parafacials bare or partially haired (hairing if present normally 
not reaching as low as bottom of eye). 9 normally without proclinate orbital setae (one or 
two present in some specimens). Arista micropubescent. Humeral callus with 3-4 setae, 
inner one or two sometimes scarcely differentiated from hair, very raiely only outer two present. 
Posthumeral setae absent or one inner posthumeral developed just mesad of humeral callus. 
Usually one post ia seta, occasionally none, rarely a small second seta developed. Scutum 
with or without some supernumerary prescutellar setae. Postalar callus with 4 (rarely 5) 
strong setae. Suprasquamal ridge thickly long haired. Scutellum flattened; with 4-10 pairs 
of marginal setae (these sometimes, stiff, straight, slightly spiniform); marginals not preceded 
by any preapical setae. Hair of lower part of pteropleuron not developed in front of level of 
posterior sternopleural seta. One sternopleural seta (o + 1), rarely trace of a small anterior 
stpl seta (especially in 9). Prosternum bare; prosternal membrane with or without some long 
soft hair. Hind tibia with long regular anterodorsal fringe (without evident ad setae amongst 
it) and without fd setae or with only one such seta (very rarely trace of second fd). Abdomen 
with median depression in last tergite (T5). T3 without median marginal setae (rarely one pair 
in 9), with lateral marginal setae. T5 with median transverse row of strong erect setae. 
Sternite 5 of § with simple non-prominent rounded lobes. ¢ genitalia with distal membranous 
part of distiphallus subequal in length to or at most only slightly longer than sclerotized proximal 
part, and with extremely large foliaceous surstyli (Text-figs 66-71). [Mainly large forms mea- 
suring 15-23 mm, with dark head ground colour and usually dark unicolorous body, never 
with bold spot pattern.] 
DISTRIBUTION. Mainly distributed throughout Australia and Tasmania, but a 
few species also in Solomon Islands, New Hebrides, Fiji and Samoa. Unknown 
from the Oriental Region and New Guinea. 
