

REVISIONARY CLASSIFICATION OF RUTILIINI 83 
most distinctive, and no other Rutiliine is known in which a surstylus of this shape 
occurs (Text-fig. 56) or in which the genitalia have strong setae with divided apices. 
Subgenus GRAPHOLOSTYLUM Macquart stat. n. 
Grapholostylum Macquart, 1851 : 196 (223). Type-species: Grapholostylum dorsomaculatum 
Macquart, 1851, by monotypy. 
Agalmia Enderlein, 1936 : 433. Type-species: Rutilia albopicta Thomson, 1869 [=Grapholo- 
stylum dorsomaculatum Macquart, 1851], by original designation. Syn.n. [Junior homonym 
preoccupied by Agalmia Enderlein, 1934 (Muscidae)]. 
Diacnosis. Parafrontals pollinose, not metallic. Epistome and genae not metallic. Facial 
carina slightly flattened to strongly convex on anterior surface, outline convex in profile and 
sharply distinguished from lunula; epistome prominent. Parafacials bare or haired. @ pro- 
clinate orbital setae usually absent, at most one pair (sometimes present one side and absent 
the other). Arista long-pubescent to short-plumose. Humeral callus with 4 setae, usually 
well developed. Posthumeral setae one or two. One very strong post ia seta [one specimen 
seen with small second fost ia]. Scutum without supernumerary prescutellar setae. Postalar 
callus with three strong setae. Suprasquamal ridge haired. Scutellum convex dorsally without 
flattening towards apex; 4-6 pairs of marginal setae; with row of well developed preapical 
setae before the marginals. Pteropleuron not haired in front of level of posterior sternopleural 
seta. Two strong sternopleural setae (1 + 1) [one specimen seen with small second anterior 
stpl]. Prosternum bare; prosternal membrane bare or sparsely haired (variable in same species). 
Hind tibia without anterodorsal fringe, usually two distinct ad setae and two or three pd setae. 
Last abdominal tergite without median depression, evenly convex across its width and sides 
strongly tapering posteriorly (hypopygium prominent and well visible in profile). T3 without 
median marginal setae or with one pair or with a transverse row of semi-erect small median 
marginals developed to variable extent; T3 with lateral marginal setae. T5 with some long 
fine erect discal setae (often irregular in § but tending to form transverse preapical row in 9). 
Sternite 5 of ¢ acuminate posterolaterally and with a submedian pair of conspicuous downwardly 
directed prominences (Text-fig. 33), these visible in situ in profile. ¢ genitalia with apical 
membranous part of distiphallus whip-like, exceptionally long and slender (about twice as long 
as sclerotized basal part of distiphallus: Text-fig. 38), surstyli long and narrow and cerci longer 
than surstyli (Text-fig. 64). [Medium-sized to large species with white pollinose spots over 
mesopleura and sternopleura and often with white pollinose spots distinguishable over humeral 
calli to notopleura and in supra-alar areas]. 
DistRIBuTION. Eastern Australia from Tasmania to Queensland. 
Discussion. Gyvapholostylum was originally described by Macquart (1851) as a 
genus allied to Rutilia having one included species, viz. G. dorsomaculatum. For 
many years the identity of Macquart’s genus remained enigmatic, but Townsend 
(1932 : 38; 1936 : 153; 1938 : 416) rightly showed that Grapholostylum is a true 
Rutiliine. Regrettably, Enderlein (1936 : 441), guessing from the description of 
Macquart and overlooking Townsend’s (1932) paper, decided that the genus belonged 
to the Ameniinae and placed it near Amenia Robineau-Desvoidy—a rather bad case 
of misidentification, which was corrected by Crosskey (1965 : 103-106), who showed 
that the misidentified G. dorsomaculatum sensu Enderlein is actually Amenia sex- 
punctata Malloch. For the present work Macquart’s type-material of G. dorso- 
maculatum (including the lectotype designated elsewhere: Crosskey (1971 : 271)) 
has been studied in detail and Townsend’s placement in the Rutiliini found to be 
correct ; it should be emphasized, however, that the name applies to a valid species 
