REVISIONARY CLASSIFICATION OF RUTILIINI 95 
ultimately be found that possess adventitious discals). But presence or absence of 
discals on T3 and T4 is a useful rule-of-thumb distinction between Amphibolia and 
Rutilia. Those Rutilia (a few species) which have the suprasquamal ridge bare are 
immediately separable from Amphibolia by this character (all known species of 
Amphbolia have the suprasquamal ridge fully haired). Likewise those Rutilia 
species that have a very broad medially depressed T5 are at once separable from 
Amphibolia, which does not have a deep median depression in this tergite (the last 
tergite in all Amphibolia species is truncate subconical with scarcely any trace of 
flattening or hollowing medially at the tip). 
The two subgenera of Amphibolia are distinguished by the following key: 
KEY TO THE SUBGENERA OF AMPHIBOLIA 
t Thorax and abdomen with bold black-and-white pattern. Lobes of fifth sternite of 
3 simple. ¢ hypopygium with surstyli shorter than cerci (Text-figs 87-89). 
Apical pair of scutellar setae odes as strong as other scutellar marginal 
setae . . AMPHIBOLIA Macquart (p. 95) 
- Thorax and abdomen without such pattern. Lobes of fifth sternite of g with small 
tooth or prong on inner edge near apex (Text-figs 34 & 35). dg hypopygium with 
surstyli longer than cerci (Text-figs 85 & 86). Apical pair of scutellar setae much 
weaker than other marginal setae (sometimes absent) 
PARAMPHIBOLIA Brauer & Bergenstamm (p. 100) 
Subgenus AMPHIBOLIA Macquart 
Amphibolia Macquart, 1843 : 278 (121). Type-species: Amphibolia valentina Macquart, 1843 
by original designation and monotypy. 
Diacnosis. Thorax black with bold discrete white-pollinose spots on dorsum. Calyptrae 
_ black or brownish black. Scutellum with apical pair of setae subequal in size to other marginal 
setae. Abdomen conspicuously patterned in contrasting black and white-pollinose araes. 
Thoracic and abdominal chaetotaxy not spiniform. Lobes of g fifth sternite simple, without 
subapical tooth oninneredge. ¢g hypopygium with short broad surstyli (often slightly bifurcate) 
which are shorter than cerci. 
DISTRIBUTION. Australia and Tasmania, Lord Howe Island, New Guinea. 
_ Discussion. This subgenus includes some of the most attractive and beautifully 
marked flies among the whole of the Rutiliini which are at once recognized by eye 
_ from their bold black-and-white coloration of the thorax and abdomen combined 
with their bright yellow heads. Not all species, however, which belong in the 
_ subgenus are quite so distinctive, for some have the white areas less boldly displayed 
than others, and not all of them have the yellow head colour. In general the species 
fall into two groups, one including the type-species (valentina) in which the head is 
mainly yellow or at most reddish brown and the abdomen has (as part of the pattern) 
discrete rounded black spots, and another including species in which the head 
ground colour is distinctly black and in which the abdomen (though possessing a 
' pattern) lacks definite isolated black spots; these two groups, which are both Austra- 
lian, are however interconnected by a new species (papuana) here described from 
New Guinea, which has the black ground colour of the head but also has discrete 

