

REVISIONARY CLASSIFICATION OF RUTILIINI 107 
life-history between Chrysopasta and termites: Townsend (1936 : 152), referring to 
Malloch’s note, wrote that ‘Echrysopasta has been reared in West Australia from a 
puparium found in nest of Eutermes, but it is likely that the maggot was a white 
grub parasite whose host in the soil had accidentally become incorporated into the 
termite nest’. This is a reasonable supposition, and the reference to a possible 
inquiline relationship between Chrysopasta and termites made by Colless & McAlpine 
(1970) on page 737 of The Insects of Australia must be treated with caution. 
INCLUDED SPECIES 
Chrysopasta elegans (Macquart). AUusTRALIA (Western Australia). [Holotype 
examined]. 
versicolor Brauer & Bergenstamm. 
zabirna (Walker). [Lectotype examined]. 
Genus PRODIAPHANIA Townsend 
Diaphania Macquart, 1843 : 277 (120). Type-species: Diaphania testacea Macquart, 1843, 
by monotypy. [Junior homonym preoccupied by Diaphania Hiibner, 1818 (Lepidoptera).] 
Prodiaphania Townsend, 1927: 159. Replacement name for Diaphania Macquart, 1843, 
preoccupied by Diaphania Hiibner. Type-species: Diaphania testacea Macquart, 1843, by 
automatic fixation. 
[Senostoma sensu authors, not Macquart. Misidentification.] 
Diacnosis. Facial carina broad and flattened, sides subparallel or only slightly convergent 
below, widely separating antennal bases. Epistome strongly prominent, subnasute, face in 
profile deeply excavate between epistome and carina. Head of ¢ not holoptic but frons strongly 
narrowed, not wider than facial carina, upper eye facets not enlarged. Parafacials haired on 
upper parts. Buccal opening elongate and strongly narrowed medially, at narrowest point 
subequal in width to or even a little narrower than facial carina. Genal dilation well developed. 
Head pollinose, non-metallic. Arista bushy plumose or long-pubescent. Palpi exceptionally 
small, sometimes papilliform, shorter than third antennal segment. Proboscis with mentum 
parallel-sided or at most only a little tapering in profile. Prosternum and prosternal membrane 
bare. Scutellum with apical pair of setae inserted conspicuously lower than other marginal 
setae; total of four to nine pairs of scutellar marginals; disc of scutellum convex. Postalar 
callus with four to six strong setae (three in holotype of cygnus). Postalar wall bare. Supra- 
squamal ridge bare. Upper calypter extraordinarily developed, subequal in length in both 
sexes to lower calypter. Tegula with normal long wiry posterior setulae. Costal base very 
strongly explanate (wings in resting position appearing to have prominent ‘shoulders’). Ab- 
dominal venter without strongly developed setae on tergites; T3 dorsally without median 
marginal setae or at most with one pair in 2, marginals of T4 short; no abdominal discal setae. 
T5 truncate subconical, without median depression (at most with only slight flattening or 
hollowing at extreme tip). 
DIsTRIBUTION. Australia only; occurring from Western Australia to Victoria and 
north to Queensland, present in Tasmania. 
Discussion. There has been some confusion in the nomenclature of this distinc- 
tive genus, to which it is necessary to refer before considering the characteristics and 
affinities. Macquart’s name Diaphania is preoccupied by Hiibner’s use of this 
name in Lepidoptera, but Brauer & Bergenstamm (1889 : 126; 1893 : 175) and 
Engel (1925 : 343)—being evidently unaware of the homonymy—continued to 
use Diaphania Macquart as a valid name; later, however, Townsend (1927 : 159) 
