TACHINIDAE OF AUSTRALIA 59 
Tribe GLAUROCARINI 
This is a very small tribe that is confined to the Old World tropics and subtropics 
and contains only the two genera Glaurocara Thomson and Doddiana Curran; 
other supposed Old World genera have been described (Oestrocharis Villeneuve, 
Semisuturia Malloch and Oestrocara Townsend) but these are now treated as 
synonyms (Crosskey, 1962). Townsend (1936) placed several New World genera 
in the tribe but none of these have the fused abdominal tergites of true Glaurocarini 
and evidently do not belong. Doddiana occurs from Java to Queensland and 
New South Wales, and Glaurocara occurs in Malaya, the south-east Asian archipelago, 
the Ethiopian Region and Mauritius; Mauritius is the type-locality of G. flava 
(type-species of Glaurocara) and is also a locality into which Doddiana mellea 
(Wiedemann) has been unsuccessfully introduced from Java for control of sugar-cane 
moth borer. 
The only host recorded for the genus Doddiana is the sugar-cane moth borer 
Chilo sacchariphagus (Bojer) (Lepidoptera: Pyralidae), which is apparently attacked 
by Doddiana mellea in Java. This is a curious record, as Doddiana is without 
doubt very closely allied to Glaurocara and the type-species of this genus, viz. 
G. flava Thomson from Africa, is a parasite of nocturnally active tettigoniid 
Orthoptera (Crosskey, 19655). It is odd that species of two such similar genera 
as Doddiana and Glaurocara should attack hosts as different as a pyralid and a 
bush-cricket. It seems probable, however, that Orthoptera are the normal hosts 
of the Glaurocarini, as Glaurocara first-stage larva is a perfect planidium closely 
similar to that of the Ormiini and the ormiines (so far as is known) are exclusively 
parasites of Grylloidea and Tettigonioidea. Glawrocara also resembles the Ormiini 
in being nocturnally active and in having the pallid luteous yellow coloration 
characteristic of night-flying Diptera, and its female reproductive system is of 
Townsend’s type XXXII which is elsewhere found only in the Ormiini. On these 
grounds it appears that the Glaurocarini are closely allied to the Ormiini, although 
they differ obviously from the latter tribe by having the sutures between the 
abdominal segments largely fused above and in having a normal prosternum (without 
trace of the inflation of the prosternal region as occurs in Ormiini). 
The chief characteristics of the tribe are as follows. Head unusually small for body size; 
eyes usually with some very minute sparse hairing (appearing bare at first glance); ¢ head 
with eyes strongly approximated and frons reduced but not holoptic; ocelli present, usually 
rather raised; inner vertical setae present in both sexes, usually convergent or crossed; ¢ 
without proclinate orbital setae, 9 with two pairs (very strong); 9 with one pair of reclinate 
orbital setae; parafacials bare; frontal setae moderately strong, rows reaching to a level with 
first antennal segment; epistome flat, invisible in profile, face rather flat; oral cavity, proboscis 
and palpi very small; vibrissae present, inserted about level with or just above epistomal 
margin; facial ridges with a few setulae above vibrissae, otherwise bare; genal dilation weak, 
only reaching at most half way forwards on the genal region, usually bearing some rather 
strong setae or setulae (Text-fig. 34); occiput moderately flat above; antennae small (length 
about half the eye height), inserted about on a level with eye middle; arista with short or 
long pubescence; prosternum not inflated, bare, prosternal membrane bare; propleuron bare; 
prostigmatic and propleural setae very strong; humeral callus with two strong setae (sometimes 
a weak third seta present mesad of main two); mesonotal chaetotaxy usually as in Text-fig. 57; 
o + 2 or o + 3 ia setae; 2 + 3 dc setae; acy setae variable; pra seta weak or absent, second 
