88 RR. W. CROSSE 
— Inner vertical setae subparallel. Abdominal Tr + 2 and T3 each with one pair of 
median marginal setae (those of Ti + 2 sometimes hair-like or virtually absent). 
Abdomen usually with one or two distinct discal setae on one or both intermediate 
tergites. Humeral callus with three main setae standing either in a straight 
line or in a triangle. Q without outwardly directed prevertical setae and with 
the usual pair of outer vertical setae. Pre-alar seta very small, shorter than 
first post ia seta . 3 : : ¢ 
2 Humeral callus with the three main setae Sear ina straight ae. Lower calypter 
with the outer edge distinctly bent downwards. Arista with second segment 
elongate, about 4—5 times as long as broad. Head with two pairs of strong 
reclinate orbital setae. Mid tibia with one ad seta. Vibrissae level with 
epistomal margin : : ‘ ETHILLA Robineau-Desvoidy 
— Humeral callus with the ee main ‘sete eden in a distinct triangle. Lower 
calypter not noticeably bent downwards. Arista with second segment not 
elongate, only about as long as broad. Head without definite reclinate orbital 
setae (at least in g, probably one pair in 9). Mid tibia with two ad setae. 
Vibrissae inserted well above level of epistomal margin 
MYCTEROMYIELLA Mesnil 
nN 
Tribe WINTHEMIINI 
This group is very closely allied to the Carceliini and the Sturmiuini and its 
recognition as a tribe is only doubtfully justified. Townsend (1941) placed Winthemia 
and its allies in the Sturmiini, but recent workers universally treat the winthemiines 
as a valid tribe and this course is followed for the present work. The arrangement 
of the humeral setae in Winthemia is rather distinctive, there being (in males at 
least) five setae in which three are strong and are spaced in a subtriangular arrange- 
ment; in this respect Winthemia differs from the Sturmiini in the strict sense in 
which the humerals are four arranged as three in a straight line and one set forwards. 
Mesnil (1944) used the presence of five humeral setae as a key character for recognition 
of the winthemiines, but in reality the character is poor — often in other winthemiine 
genera than Winthemia itself (e.g. in Nemorilla) five setae may not be definitely 
developed in males and are typically not developed in females (which usually have 
four humerals). 
The winthemiines are best recognized among the Australian Goniinae by the following 
features in combination (though some Sturmiini and some Carceliini may show some of the 
features together): eyes densely haired; males without definite reclinate orbital setae; hind 
tibia (especially in males) with a well developed close-set ad fringe; barette completely haired 
(or nearly so); scutellum with strong crossed horizontal apical setae and with subapical setae 
inserted widely apart (their bases as distant from each other as base of either from the basal 
seta of the same side, or even more distantly separated than this) (as in Carcelia, Text-fig. 72). 
A noteworthy feature of several forms is the presence of pale yellow hair on the prosternum 
instead of the usual black hairs or setulae. 
There is close superficial resemblance between Winthemiini and Ethillini, and 
as the genus M ycteromyiella (Ethillini) could easily be confused with the winthemiines 
it is included in the following key. 
Mesnil (19494 : 80) treated Crypsina as a subgenus of Winthemia but it is here 
preferred to revert to its original generic status. This course maintains Winthemia 
as a more homogeneous genus, though — certainly — Crypsina is very closely allied. 

