go RW. CROSSKE ¥ 
2 Eyes bare. Apical scutellar setae very small, directed upwards and either not 
meeting or crossing at tips. Subapical scutellar setae not conspicuously distant 
from each other, their bases separated by a distance not greater than that 
between a subapical seta and its corresponding basal seta. Upper occiput 
with some fine black setulae behind postocular row 
ARGYROPHYLAX Brauer & Bergenstamm 
— Eyes haired. Apical scutellar setae strong, crossed and horizontal. Subapical 
scutellar setae very widely separated, distance between bases much greater 
than that between base of a subapical seta and its corresponding basal seta 
(Text-fig. 72). Upper occiput without black setulae behind postocular row ; 3 
3 Dorsocentral setae 3 + 4. Scutellum with extremely strong apical setae that are 
at least as strong as, and usually larger than, the lateral setae. Hairing of 
eyes long and dense, very conspicuous. Barette normally with hairing, at 
least at anteriorend. Wingshyaline. Not conspicuously black forms : 
— Dorsocentral setae 3 + 3 (four post dc setae in occasional specimens). Scutellum 
with apical setae normally shorter and weaker than lateral setae. Hairing of 
eyes rather short and usually rather sparse, not so immediately conspicuous. 
Barette totally bare. Wings suffused with brown anterobasally. Conspicuously 
black forms with shining abdomen which shows bright silver pollinose areas on 
each side of T3-Ts5 (these especially noticeable on T4, at least in Q) 
ARGYROTHELAIRA Townsend 
4 Facial ridges bare. Barette when haired with hairing confined to anterior half 
(except sometimes in specimens of Carcelia s. str.) . CARCELIA Robineau-Desvoidy 
— Facial ridges closely setulose up most of their height (Text-fig. 48). Barette 
completely haired : ‘ : : : : : CARCELIMYIA Mesnil 
KEY TO AUSTRALIAN SUBGENERA OF CARCELIA 
1 Mid tibia with a submedian v seta. Ocellar setae absent or very small, fine and 
wiry. 4 with one pair of reclinate orbital setae, 9 usually with two pairs (specimens 
with two such setae on one side and one on the other occur). Hind coxa bare 
or setulose posterodorsally . c subgenus CARCELIA Robineau-Desvoidy 
— Mid tibia without a v seta. Ocellar setae very strong (except in muvina where 
absent or hair-like). Both sexes with two pairs of reclinate orbital setae. Hind 
coxa entirely bare posterodorsally. : . subgenus SENOMETOPIA Macquart 
Tribe ANACAMPTOMYIINI 
Townsend (1940) recognized a tribe for Anacamptomyia and its allies and 
placed it near Compsilurini (i.e. Blondeliini), but Mesnil (1944) included the 
anacamptomyiines among the carceliines. On adult morphology Mesnil’s placement 
seems to be more appropriate than Townsend’s, and in fact it is difficult to find 
characters that satisfactorily differentiate the Anacamptomyiini from the Carceliini. 
Nevertheless the Anacamptomyiini is here treated as a valid tribe because of its 
biology: the members of the tribe are unique amongst the Goniinae in being 
parasites of paper-making wasps (such as Belonogaster and Polistes), and this 
host-relationship is so exceptional that it appears justified to give it considerable 
weight in assessing whether or not to treat the flies involved as carceliines or not. 
It appears possible that the resemblances between Carceliini and Anacamptomyiini 
are mainly convergent. 
