OF PSEUDOCOCCIDAE & ERIOCOCCIDAE 5 



Davies for reading part of the manuscript, and highly valued criticism. The 

 technical remarks received from Dr. J. H. Giliomee, the earlier Ph.D. student in the 

 department, are thankfully acknowledged. My deep appreciation is also due to 

 the Trustees of the British Museum (Nat. Hist.), and also to Dr. D. J. Williams, 

 of the Commonwealth Institute of Entomology, for providing specimens of four 

 species ; a large number of other workers have also kindly furnished material from 

 their collections, or even took the trouble to collect and breed the males specially 

 for the purpose of this study, namely : the late Prof. N. S. Borchsenius, of the 

 Zoological Institute, Academy of Sciences, Leningrad ; Dr. G. M. Das, of the 

 Tocklai Experiment Station, Cinnamara, Assam, India ; Mr. G. De Lotto, of 

 the Research Institute for Plant Protection, Pretoria ; Mgr. A. Dziedzicka, of the 

 Pedagogical Higher School, Krakow, Poland ; Mr. A. I. Ezz, of the Department of 

 Entomology, Ministry of Agriculture, Cairo, U.A.R. ; Mr. J. M. Hoy, of the 

 Entomology Division, Dept. of Scientific and Industrial Research, New Zealand ; 

 Prof. M. Kosztarab, Virginia Polytechnic Institute, Blacksburg, Virginia ; Dr. G. 

 Matesova, of the Institute of Zoology, Academy of Sciences, Alma-Ata, Kazakhstan ; 

 Mr. D. N. McNutt, of the Agricultural Department, Mbale, Uganda, East Africa ; 

 Ing. N. Mitic-Muzina, of the Institute for Plant Protection, Belgrade ; Mr. J. 

 Munting, of the Plant Protection Research Institute, Pretoria. 



In this opportunity, I would like also to express my gratitude to Prof. Dr. Y. M. 

 Ezzat, Head of the Department of Zoology, University of Assiout, U.A.R., for his 

 constant and unfailing encouragement, which has always given me a great deal 

 of confidence. 



The Ministry of Agriculture of the United Arab Republic (Egypt) has the credit 

 of originally awarding me a study leave, which was later elevated to a full mission ; 

 a financial support without which this work would have not been accomplished. 



INTRODUCTION 



This work is the fourth in a series of detailed morphological and taxonomical 

 studies on the males of Coccoidea (Theron, 1958, studied the general morphology of 

 7 species representing 4 families ; Ghauri, 1962, described in detail 24 species of the 

 family Diaspididae ; Giliomee, 1967, similarly treated 23 species of the family 

 Coccidae) carried out by the postgraduate students in the Department of Zoology 

 and Applied Entomology, Imperial College, suggested and supervised by Dr. K. 

 Boratynski. The present publication deals with 24 species belonging to two groups, 

 the pseudococcids (17 species) and the eriococcids (7 species), whose taxonomic 

 status is still controversial ; they have been considered as (1) merely tribes of one 

 subfamily, Pseudococcidae (Cockerell, 1899 ; Balachowsky, 1948) ; (2) subfamilies 

 of one family (Balachowsky, 1942 ; Borchsenius, 1949), or (3) separate families, 

 with rather remote connection between them (Ferris, 1937, 1957 ; Borchsenius, 

 1963). Apart from the grouping of certain pseudococcid genera by some authors 

 (the Rhizoecus group including 8 genera by Hambleton, 1946, and the tribe 

 Planococcini by Ezzat & McConnel, 1956), no comprehensive classification of the 

 pseudococcids or the eriococcids has been hitherto proposed. The purpose of the 



