io M. W. NIELSON 



Dr Charles H. Triplehorn, Ohio State University, Columbus (OSU); Dr S. L. Tuxen, 

 Universitetets Zoologiske Museum, Copenhagen, Denmark (UZM); Dr George 

 Wallace, Carnegie Museum, Pittsburgh (CM); Dr Richard G. Wilkey, California 

 State Department of Agriculture, Sacramento (CSDA); Dr Keizo Yasumatsu, 

 Kyushu University, Fukuoka, Japan (KU); Dr David A. Young, North Carolina 

 State University, Raleigh (NCSU). 



Subfamily COELIDIINAE Dohrn 

 Coelididae Dohrn, 1859 : 84. Type-genus: Coelidia Germar, 1821. 



Concepts of the subfamily Coelidiinae proposed by earlier workers (Matsumura, 

 1914; Oman, 1936; 1949; Evans, 1947; 1971; Linnavuori, 1959; 1960a; 19606; 

 Metcalf, 1964) have been based for the most part on studies of a few genera within 

 certain geographical areas. Earlier attempts to fully define the generic affinities 

 of Coelidiinae were difficult since up until this revision the generic concept of 

 Coelidia per se never had been accurately assessed, thus previous concepts formu- 

 lated were actual assumptions based on presumed species of Coelidia without 

 recourse to the type-species of Coelidia. 



The syntypes of the type-species, Coelidia venosa Germar were recently found 

 in the Naturhistorisches Museum in Vienna, along with the types of three other 

 Germar species, pruinosa, poecilia, and variegata, all described in the genus Coelidia. 

 Coelidia venosa is represented by two female specimens, one of which I shall be 

 designating as a lectotype in my treatment of that species in the tribe Coelidiini. 

 This specimen has been compared to and accurately associated with a male 

 specimen on loan from the American Museum of Natural History, New York. 

 The true identity of venosa and its generic characteristics thus establishes a concept 

 within which all other related genera must fall so that an appropriate definition 

 of the subfamily can be made. 



When Matsumura (1914) treated the Coelidiinae of Japan using the German 

 vernacular, Coelidinen, he actually used the characteristics of Coelidia conspersa 

 (Stal) to form his own generic concept of Coelidia in his treatment of nine species 

 of the Japanese fauna. None of these species belong to Coelidia but have been 

 relegated to two new genera and one old genus in my treatment of the tribe 

 Coelidiini. 



Kirkaldy's (1901) and Oman's (1936) selection of Coelidia venosa Germar as the 

 type-species of the genus was presumably based on priority by pagination rather 

 than as being 'represential' of the genus since they did not have access to the 

 type-specimen. 



Evans (1947) was first to broaden the generic affinities of the subfamily when he 

 treated 32 generic names, 26 of which he considered valid. He characterized 

 the group principally on gross external morphology, with emphasis on head 

 characters and venation of the elytra. Genera that were included varied from 

 those having incomplete tegminal venation, i.e., both M 1+2 and cross vein m — Cu 3 

 wanting (one closed anteapical cell) to those having complete venation (three 



