REVISION OF COELIDIINAE n 



closed anteapical cells). It is noteworthy that all genera with complete tegminal 

 venation have an intact ninth sternum (male valve) which is a separate, ventral 

 triangular sclerite attached laterally to the pygofer. On the other hand, all genera 

 with incomplete tegminal venation (vein Afj _ 2 missing or one closed anteapical 

 cell) have the male valve fused ventrally to the pygofer. I consider the combination 

 of these latter characters as well as other characters discussed below as significant 

 subfamily attributes and therefore have excluded all genera from the subfamily 

 Coelidiinae treated by Evans (1947; 1971) that do not possess these characters. 

 The intermediate genera (those with two closed anteapical cells and separate male 

 valve) have also been excluded. 



The following genera are excluded with provisional assignment to other sub- 

 families: Aeturnus Distant, Selenopsis Spinola and Wadktipfia Linnavuroi, to 

 the Deltocephalinae; Dardania Stal, Iberia Kirkaldy and Stegelytra Mulsant & 

 Rey to the Stegelytrinae. The genera A letta Metcalf, Alocoelidia Evans, Caelidioides 

 Signoret, Equeefa Distant, Imquerus Ghauri, Iturnoria Evans, Malagasiella Evans, 

 Palicus Stal, Protonesis Spinola, Cyrta Melichar, Doda Distant, Kasinella Evans, 

 Kunasia Distant, Placidellus Evans, Placidus Distant, Sabimamorpha Schumacher 

 and Toba Schmidt cannot be assigned to any other existing subfamily; new 

 subfamilies will be proposed for them in forthcoming studies. 



Oman (1949) was first to assess the significance of genitalia of Coelidiinae in 

 his descriptions of two nearctic genera (Coelidia s. 1. and Tinobregmns) and establish- 

 ment of the tribe Tinobregmini. 



Linnavuori (1959) presented perhaps the most complete concept of the 

 Coelidiinae in his characterization of the group. His concept was no doubt 

 influenced by a prior work (Linnavuori, 1956) on descriptions of 21 new species 

 and redescriptions of 13 old species of Coelidiinae from the Neotropical realm. 

 Linnavuori (1959) utilized both external characters of the body and the internal 

 male genitalia. However, in his studies of the Coelidiinae of Fiji Islands (1960a) 

 and Micronesia (19606) he apparently followed Wagner's (1951) system of hierarchical 

 classification. The Coelidiinae were treated under a subfamily group, the 

 Cicadellides, characterized as those subfamilies possessing ocelli on the anterior 

 margin of the head or crown, valve usually more or less triangular, and articulated 

 with pygofer (fused to pygofer in primitive forms) and plates usually triangular. 

 Aside from the Typhlocybides, the other subfamily group, Iassides, are defined as 

 subfamilies having ocelli distinctly on the face or the head, the ninth sternum of 

 the male never triangular and always fused to the pygofer, and the male plates 

 always parallel sided. 



The Coelidiinae as presently constituted do not fit into either group. Among 

 6 tribes, nearly 100 genera and over 600 species treated in my revision, the subfamily 

 characters include major attributes common to both subfamily groups. These 

 are: ocelli always near the anterior margin of the crown or laterally in long-headed 

 forms, male valve always fused ventrally to the pygofer, and the male plate elongate, 

 parallel sided, sometimes laterally appressed, and never triangulate. 



Wagner's hierarchical system of classification has some notable limitations, 

 and while it is not my purpose in this paper to propose changes that are necessary, 



