TACHINIDAE OF ORIENTAL REGION 



'7 



all Phasiinae have bare eyes, and any specimen showing hairy eyes must belong 

 elsewhere, whilst if a specimen is known to have had a hemipterous host then it 

 must belong (on present evidence at least) to the Phasiinae. 



Four tribes are here recognized in the Phasiinae, the Phasiini, Cylindromyiini, 

 Leucostomatini and Eutherini, in accordance with the current trend of specialists 

 to reduce the number of tribal entities to be granted validity. Some authors 

 have treated, or continue to treat, Trichopoda Berthold and its allies as the tribe 

 Trichopodini, and Gymnosoma Meigen and its allies as the tribe Gymnosomatini 

 (both tribes being placed close to Phasiini) and I have myself treated the Trichopo- 

 dini as a valid tribe in a recent work (Crosskey, 1973ft); but it is now plain that if 

 the world fauna is considered as a whole there is no real character gap that justifies 

 the tribal segregation of the Trichopoda and Gymnoscma complexes from typical 

 Phasiini, and the trichopodines and gymnosomatines are here regarded as an integral 

 part of the Phasiini. The existence of tropical genera such as Perigymnosoma 

 Villeneuve, which is intermediate between Gymnosoma and typical Phasiini, and 

 Pentatomophaga de Meijere and Bogosia Rondani, that to a considerable extent 

 interconnect Trichopoda with typical phasiines such as Ectophasia Townsend, 

 supports the case for widening the concept of Phasiini and abandoning the rather 

 valueless tribal concepts of Gymnosomatini and Trichopodini. Even so, it is not 

 possible to differentiate an enlarged Phasiini from the Cylindromyiini in a completely 

 convincing way, and the external adult characters that have to be used to distinguish 

 these tribes in the Oriental fauna - and indeed to separate all the four tribes - are 

 not very positive when they have to be crystallized into key couplets. 



Key to Oriental Tribes of PHASIINAE 



1 Face formed into a broad swollen carina that extends from the antennae to the 



epistome and has a sharp median edge (e.g. as Text-fig. 27). Wing boldly patterned 

 with two dark brown cross-bands (broad submedian band and narrow subapical 

 band, Text-fig. 90) and with unusually elongate blackish brown alula. Usually 

 three post ia setae (if only two differentiated then foremost one closer to the hind 

 one than to the transverse suture). Four or five post dc setae (often rather fine) 



EUTHERINI (p. 34) 

 - Face not carinate or if slightly swollen medially not formed into a sharp edge. Wings 

 and alula not so. Fewer than three post ia setae, none, one or two (if two then 

 the foremost one as close to or closer to the transverse suture than to the hind one). 

 From one to three post dc setae, or if exceptionally four then usually some of them 

 almost hair-like ............ 2 



2 Abdominal vestiture entirely or almost entirely hair-like (sometimes the tergite 



marginal hairing a little stronger than the rest). Abdomen with fully exposed 

 sternites and usually with dorsoventrally flattened or subglobular shape. One 

 or two post dc setae on the posterior half of the scutum (rarely one or two tiny 

 hair-like dc in addition). One post ia seta or none. Pre-alar seta absent (except 

 in Gymnosoma) and second sa seta absent (supra-alar region of the scutum therefore 

 normally with only one seta, the first sa). Lower calypter broad, nearly straight 

 or very slightly concave on its posterior edge (except enormous kidney-shaped 

 flap in Compsoptesis). Femora sometimes with ventral comb-like rows of strong 

 spinules. Posteroventral declivity of the thorax usually widely membranous 

 medially and metacoxae and abdominal base not widely separated. Bend of vein 



