TACHINIDAE OF ORIENTAL REGION 33 



postabdomen with sternite 6 -f- 7 in the form of a simple transverse plate bearing 

 a very few inconspicuous posterolateral spines, and with a few. similar spines on 

 the posteroventral corners of T6 + 7 . . . . . epalpata Townsend 



[The $ holotype of this species is aberrant, the right palp being completely 

 absent and the left palp represented by a small papilla. Other characters, 

 including the postabdomen, confirm the identity as being with a species that 

 normally possesses well developed clubbed palpi and confirm the indisputable 

 assignment of epalpata to Lophosia and not to the Cylindromyia group of genera 

 in which palpi are lacking. It is unfortunate that Townsend did not recognize 

 the teratological state of the palpi of epalpata holotype, as this specific name is 

 scarcely appropriate to a species that normally possesses large palpi.] 



Tribe LEUCOSTOMATINI 



The external adult characteristics of this tribe have been given in an earlier 

 paper (Crosskey, 1973ft : 38). Only two named species are recorded from the 

 Oriental Region, each being the type-species of an endemic genus, but a third 

 species is known to occur in the area (a specimen from China being present in the 

 BMNH collection that does not belong in either of the endemic genera). Hennig 

 (1941 : 189) recorded ' Paradionaea orientalis Baranoff' from Formosa but this 

 name remains a nomen nudum (Sabrosky & Crosskey, 1969 : 57); the specimen to 

 which it appertains is in the DEI collection but has not been seen and may be 

 conspecific with the above-mentioned BMNH specimen from China. At least two 

 undescribed species belonging to, or close to, Dionaea Robineau-Desvoidy occur 

 in New Guinea, and one of these appears to be the same species as the undetermined 

 specimen from China alluded to. 



The position of the genus Pseudobrullaea within the Phasiinae is problematical. 

 The genus is known only from the female holotype of its type-species and is here 

 placed in Leucostomatini mainly because of the horizontal forceps-like processes 

 at the tip of the abdomen (Text-hg. in). The head profile (Text-fig. 24) is more 

 like that of Cylindromyiini, and Pseudobrullaea is somewhat intermediate between 

 typical leucostomatines and cylindromyiines. 



Of special note concerning the distribution of leucostomatine genera is the 

 apparent absence of Leucostoma Meigen from the Oriental Region (to my knowledge 

 there are no records) ; this is unexpected when it is considered that Leucostoma 

 occurs widely in the remainder of the Old World, including Australia, and also 

 in North America. 



The hosts of the Oriental species are unknown, but may be presumed to belong 

 to the Hemiptera. 



Key to Oriental Genera of LEUCOSTOMATINI 



1 Scutellum with three pairs of marginal setae.* Bend of vein M obtuse and without 

 an appendix. Parafacials bare or at most with hairing at the extreme upper ends 

 continuous with the parafrontal hairing ........ 2 



* The scutellum of the holotype of Pseudobrullaea aberrans, only known specimen of Pseudobrullaea, 

 has a fourth pore very close to the subapical seta pore but three pairs of scutellar marginal setae is 

 considered to be the normal state for the genus. 



