36 R. W. CROSSKEY 



reminiscent of the Rhinophoridae (for example the lower calyptrae and the scutellar 

 bristling), and - in the absence of host-data - it is not completely certain that 

 Cylindromyiella is a tachinid, although it undoubtedly fits the Tachinidae on the 

 preponderance of its characters. Where to classify the genus among the tachinids 

 is, however, very problematical, but it probably belongs in the 'no-man's-land' 

 of forms intermediate between the Phasiinae and the Dufouriinae, or possibly to 

 the Cylindromyiini. 



When preparing the Tachinidae section (Crosskey, in press) of A Catalog of the 

 Diptera of the Oriental Region (Delfinado & Hardy, eds) I did not have the holotype 

 of C. bakeri available for study, and tentatively placed Cylindromyiella in the 

 Dufouriini, but through the kindness of Dr Curtis Sabrosky I have now been able 

 to see the specimen and to study its characters at first hand; I now conclude that 

 it is not sufficiently certain that the genus is closely related to dufouriines for it 

 to be firmly placed in Dufouriini, and Cylindromyiella is therefore left unplaced 

 in the present work. 



On balance, it is probable that Cylindromyiella is most closely related to Phasiinae, 

 but on present knowledge it cannot legitimately be assigned to any particular 

 phasiine tribe. The genus possesses a curious combination of features, some of 

 which recall one phasiine tribe and others which call to mind other tribes. The 

 wing venation, for example (in which M has only the faintest trace of forward 

 bending), is similar to that of Cinochira Zetterstedt in the Leucostomatini, but the 

 stubby spinules on the ventral surfaces of the femora resemble those of some 

 Phasiini, and the fully sclerotized closed metathorax is like that of many Cylindro- 

 myiini (though not very deep). 



More material will be needed, preferably with host information, before a firm 

 decision can be made on where to place Cylindromyiella, but in the meantime it 

 will be helpful to record the characters shown by the holotype of C. bakeri (particularly 

 as Malloch's original description needs amplification). 



Head dichoptic; eyes slightly oblique, bare; vertex about a quarter of head width; antennal 

 axis level with eye middle, frontal and facial profile lengths subequal; frons parallel-sided, 

 interfrontal area nearly twice as wide as a parafrontal; facial regions widely diverging towards 

 the epistome, facial ridges not at all prominent and invisible in profile; parafacials very narrow, 

 almost obliterated medially by near-contiguity of the eyes and facial ridges ; epistome flat 

 and invisible in profile, vibrissal angles moderately sharp; gena narrow (about one-eighth of 

 eye height) and without definite genal dilation; occipital regions swollen medially. Inner 

 vertical setae strong and converging, outer vertical setae undifferentiated; ocellar setae small 

 and proclinate; frontal setae inclinate, five pairs, rows reaching just to lunula; orbital and 

 prevertical setae entirely absent; parafrontals, parafacials and facial ridges totally bare; 

 vibrissae moderately strong but not crossing, level with epistome; postocular setulae sparse 

 and short; occipital regions bare on upper parts behind postocular row, with a few black setulae 

 near foramen and black hairing towards postbuccae. Antennae moderately heavy, first 

 segment not prominent, third segment parallel-sided and about three times as long as second 

 segment; arista long and fine, thickened only on basal quarter and bare, basal segments very 

 short; proboscis very short, palpi strongly enlarged and flattened (longer than proboscis). 

 Thoracic shape normal; propleuron, presternum and barette bare; two humeral setae; mesonotal 

 setae rather weakly differentiated [arrangement not fully certain as centre of mesonotum 

 destroyed by pin], but apparently as follows: presutural seta very weak; no prst ia seta; one 



