TACHINIDAE OF ORIENTAL REGION 71 



and that Gibsonomyia males have a holoptic head and neither prevertical nor 

 proclinate orbital setae. But the BMNH collection contains several specimens of a 

 new species of Phyllomyini (described at the end of this section as Phyllomya 

 gibsonomyioid.es sp. n.) in which the males have a holoptic head that lacks both preverti- 

 cal and proclinate orbital setae (exactly as Gibsonomyia) but in which all other male 

 characters and all female characters are those of Phyllomya. Clearly, this species 

 is somewhat intermediate between Gibsonomyia and Phyllomya, but as all characters 

 other than the male head conformation are these of Phyllomya it is described in 

 this genus and the generic definition widened accordingly. The new species is 

 from northern India, an area previously outside the known range of the genus 

 Phyllomya. 



The following are the main characteristics of the Oriental Phyllomyini. Slightly to strongly 

 elongate forms, mainly black and often with the appearance of Minthoini. Eyes bare (except 

 in Gibsonomyia annularis). £ head holoptic or dichoptic, if the latter then with prevertical 

 and proclinate orbital setae as in 9. Ocellar setae varied, from long and very fine to absent. 

 Facial ridges bare. Parafacials without setae, bare or haired (most often haired and hairing 

 sometimes very long, dense and bushy). Face flat, epistome not visible in profile. Vibrissae 

 well developed, level with or slightly above epistomal margin. Upper occiput flat or slightly 

 swollen, with many scattered black setulae behind postocular row. Gena at least a quarter of 

 eye-height, without or with a short genal dilation. Arista long-pubescent or long-plumose. 

 Proboscis short, palpi fully developed. Presternum and propleuron bare. Humeral callus 

 with two strong setae (in males sometimes also with two long fine hair-like setae differentiated 

 in addition). 2-3 prst dc setae, 2 or 3 post dc setae. Acrostichal setae absent or represented 

 by one pair of prst acr. Pre-alar seta present, strong or weak. Second sa seta weak or repre- 

 sented by a mere hair, prst ia seta absent. One strong post ia seta preceded by one or two 

 weaker post ia (first one or two sometimes hair-like). Two or three stpl setae. Pteropleural 

 seta absent. Infrasquamal setulae absent. Posteroventral declivity of thorax membranous 

 medially. Scutellum with three pairs of marginal setae (basals, subapicals and crossed apicals) 

 but basal pair sometimes very weak. § fore tarsus not enlarged or flattened (cf. typical Minth- 

 oini). Mid tibia with one or more ad setae and with submedian v seta. Hind coxa bare on 

 posterodorsal surface; hind tibia with two or more ad and two or more pd setae, with or without 

 pd preapical seta and with or without a small pv apical seta. Wing veins bare except for a 

 few setulae on basal node of ^4 + 5. Cell i? 5 open or just closed at the wing margin. Bend 

 of vein M forming a gentle evenly rounded curve near to the wing edge. Costal base with a 

 small or very large costigial seta. Second costal sector haired ventrally. Calyptrae either 

 broad and not far removed from the scutellum or small and rather rounded and widely removed 

 from the scutellum. Abdomen moderately to very strongly elongate, subovate to fusiform, 

 Ti -f- 2 not excavate and sternites concealed. Abdominal setae long, erect, T3-T5 (sometimes 

 also Ti + 2) with discal setae, all tergites with marginals, some forms with almost complete 

 transverse rows of both discals and marginals on most tergites. 



There are no host records for the Oriental members of the Phyllomyini, but they 

 possibly attack sawfly larvae (as the European species Phyllomya volvulus (Fabricius) 

 is a parasite of Tenthredinoidea). 



Key to Oriental Genera of PHYLLOMYINI 



1 Parafacials haired. Parafacial at mid-height as wide as or wider than third antennal 

 segment. Mid tibia with two or more ad setae. Fore tibia with two pv setae. 

 Pre-alar seta long and conspicuous (stronger than or as strong as the posterior 



