80 R. W. CROSSKEY 



- Antennae dark brown on most of third segment. $ interfrontal area at its narrowest 



point about four times as wide as a parafrontal. Vertex of q* about 0-29 of head- 

 width. Arista and its hairing obviously narrower than third antennal segment. 

 Abdominal pollinosity pale greyish or silvery . . . Undetermined sp. (? sp. n.) 



[Running here are two <J specimens in BMNH collection from Java that are very 

 close to sumatrense but may represent an undescribed species.] 

 5 Basicosta unicolorous yellowish orange or orange. Bend of vein M very remote from 

 wing margin and distance on vein M between m-cu and the bend very short (not 

 more than twice as long as r-m and half as long as the M 2 appendix). Ground 

 colour of the facial regions and genae entirely reddish yellow (the pale colouring 

 best seen with head viewed from below). Tibiae almost entirely reddish yellow 



M. europaea Egger 



- Basicosta black or black-brown, at most only paler yellowish posteriorly. Bend of 



vein M not strikingly remote from wing margin and distance on vein M between 

 m-cu and the bend moderately long (at least three times as long as r-m and subequal 

 to or slightly longer than the M z appendix). Ground colour of the facial regions 

 and genae very dark reddish to blackish brown (colouring best seen from below). 

 Tibiae reddish brown to blackish ...... ? Undescribed sp. 



[This species occurs in Africa as well as the Oriental Region and in the former 



area has been misidentified by past authors as M. europaea.] 



Tribe GERMARIOCHAETINI 



Mesnil (1966) first proposed this group, as the subtribe Germariochaetina, for 

 the single monotypic genus Germariochaeta Villeneuve from China. Recently 

 he (Mesnil, 19736) has characterized the group and described a second genus that 

 belongs to it (Lophosiosoma, also monotypic, and from Formosa). Until now the 

 tribe, as it is here ranked, has been known only from the two type-species from 

 China and Formosa, but three specimens in the BMNH collection (two from India 

 and one from Java) represent three previously undescribed species and serve not 

 only to expand knowledge of the characters found in the group but also to show 

 that the Germariochaetini occur widely in the Oriental Region. 



Even including the three specimens mentioned (the holotypes of the new species 

 described at the end of this section) the Germariochaetini is such a badly known 

 group that fewer than a dozen specimens are known to exist in museum collections. 

 It is probable, therefore, that the five species now known are only a small part of 

 the actual germariochaetine fauna. Nevertheless it is already obvious that the 

 group is one of great potential interest, because it contains some of the most aberrant 

 Tachinidae and quite probably attacks some unusual host group (at present the 

 hosts are unknown). 



The strongly apomorphic nature of the germariochaetines makes it difficult to 

 guess their phyletic relationships, but Verbeke's (1962&) examination of the male 

 genitalia of G. clavata Villeneuve makes it clear that they do not belong in the 

 Phasiinae despite some external resemblance to certain Cylindromyiini - which 

 some of them resemble, for instance, in having the posteroventral declivity of the 

 thorax in the form of a fully sclerotized bridge between the metacoxae and the 

 abdominal base. Instead, Verbeke placed Germariochaeta in Echinomyiinae (i.e. 

 correctly the Tachininae) and Mesnil has accepted this placement and implied 



