94 R- W. CROSSKEY 



Tribe OXYPHYLLOMYIINI 



Mesnil (1966 : 886) proposed this group (as Oxyphyllomyina) for the aberrant 

 and monotypic genus Oxyphyllomyia Villeneuve. The only material yet known 

 consists of the three female type-specimens of the type-species, 0. cordylurina, 

 from southern China. The species is aptly named, as it bears an extraordinary 

 superficial resemblance to certain species of Scathophagidae (Cordyluridae), even 

 to the extent of having exceptionally reduced calyptrae (the lower calypter being 

 no larger than the upper one). The phyletic relationships of Oxyphyllomyia are 

 very uncertain, and as the type-material was not reared there is no host information 

 to provide clues. Both Villeneuve in the original description, and Mesnil (1966) 

 by inference from his placement of Oxyphyllomyina as subtribe 37 (next in sequence 

 to his subtribe 38, Phyllomyina), considered the affinities to be with Phyllomya 

 Robineau-Desvoidy, and certainly there are resemblances between the two in 

 general body facies. It appears more probable to me, however, that the true 

 relationships are with the Leskiini, for the head facies and structure of the proboscis 

 (Text-fig. 47) are so exactly of the leskiine type and other features of the body in 

 no way contra-indicate probable derivation from leskiine-like forms. Accordingly 

 I place the Oxyphyllomyiini next to the Leskiini. Mesnil (1975a : 1349) has summar- 

 ized some of the characters of the group but a more extensive account is given 

 below. 



The characteristics of the Oxyphyllomyiini are as follows. Head shape as Text-fig. 47, 

 head nearly as long as its height and without facial carina, frontal and facial profiles of about 

 equal length, occipital and postbuccal regions strongly swollen. Eyes bare. Gena without 

 dilation, as deep as half the eye-height. Epistome very slightly warped forwards from the 

 face but not noticeably projecting in front of the vibrissal insertions. Vibrissal angles slightly 

 prominent, vibrissae well developed and about level with the epistomal margin. Inner vertical 

 setae strong and erect, outer vertical setae absent. Ocellar setae present, proclinate. Frontal 

 setae sparse, rows extending to the level of the first antennal segment. One pair of erect- 

 reclinate orbital setae, 9 with one pair of proclinate orbital setae [<$ unknown, ? proclinate 

 orbitals in this sex also]. Facial ridges bare. Parafacials very weakly haired on upper parts. 

 Setulae of postocular row strong and irregular, but very sparse; upper occiput with sparse but 

 rather strong black vestiture. Antennal axis far above the level of the eye middle, first antennal 

 segment prominent, antennae long and nearly reaching to the epistome; arista pubescent, 

 basal segments very small (no trace of elongation). Palpi fully developed, slender. Proboscis 

 very long and slender (about one and a half times as long as head height, Text-fig. 47). Pro- 

 sternum and propleuron bare. Two humeral setae (both strong), o -\- 1 ia seta, acr setae 

 absent (or ? prescutellar pair present). 2+3 strong dc setae, pra seta absent. Two strong 

 sa setae. Propleural seta present (weak), prostigmatic seta present (strong). Two stpl setae. 

 Pteropleural seta very long and strong. Hypopleural setae very weak and very few. Infra- 

 squamal setulae absent. Scutellum small, subtriangular, with only one pair of setae (very 

 strong widely divergent subapicals inserted high up on the scutellar tip, Text-fig. 72). Post- 

 scutellum almost absent, forming only a very slight convexity. Posteroventral declivity of 

 the thorax membranous medially. Wings (Text-fig. 95) long and narrow, without pattern, 

 clear. Veins (including the node) bare. Cell i? 5 open to the wing margin. Bend of vein 

 M forming an evenly rounded curve near to the wing edge; m—cu straight and meeting M 

 slightly nearer to the bend than to r-m; last section of Cu x exceptionally short (less than one- 

 third of length of m-cu). Second costal sector sparsely haired ventrally. Calyptrae very 

 small, lower calypter subequal in size to or smaller than the upper calypter, apical margin of 



