TACHINIDAE OF ORIENTAL REGION 121 



Ti + 2 and T3 (median marginals on these tergites are present in the Phorocerosoma- 

 Gynandromyia complex though they may sometimes be very weak in P. vicarium). 



The only genera other than those so far mentioned that are found in the Oriental 

 Region are Paratryphera (of which Mesnil, 1970ft : 117, has recently described a 

 species from India) and Mycteromyiella. The first of these is undoubtedly an 

 ethilline, but the tribal affinities of Mycteromyiella are uncertain and its placement 

 in Ethillini is mainly for want of anywhere more appropriate to assign it (see Crosskey, 

 19736 : 87). Several of the adult characters of Mycteromyiella, when taken in 

 conjunction with the characters of some undescribed ethilline forms from the New 

 Guinea area, suggest, however, that Mycteromyiella might be phyletically close 

 to Phorocerosoma. This possibility is supported by what is known of the orthop- 

 teroid hosts: Mycteromyiella parasitizes Phasmatodea (see Crosskey, 1968) and 

 Phorocerosoma attacks acridoid grasshoppers, the genera having an essentially 

 similar biology. No actual host records yet exist for Phorocerosoma from within 

 the Oriental Region proper, but Iwata & Nagatomi (1954) have discussed the biology 

 of P. forte Townsend (now a junior synonym of P. vicarium Walker) as a parasite 

 of Oxya japonica Willemse (now 0. yezoensis Shiraki) in Japan. 



The genus Ethilla is unknown from the Oriental Region but is likely to occur 

 there as it is known from New Guinea and Australia as well as southern Europe. 



Key to Oriental Genera of ETHILLINI 



1 Vibrissae inserted far above the level of the epistomal margin (at a distance at 



least as great as the width of the third antenna] segment). Lower calypter normal, 

 not bent downwards on its outer margin. [Forms with sturmiine facies] 



MYCTEROMYIELLA Mesnil 



- Vibrissae inserted about on a level with the epistomal margin. Lower calypter 



bent downwards on its outer margin. [Forms without a definite sturmiine facies] . 2 



2 Scutellum with three pairs of marginal setae (lateral setae absent). Inner vertical 



setae subparallel. Humeral callus with the three main setae standing in line. 

 Pre-alar setae subequal in size to, or smaller than, first post ia seta. $ without 

 outwardly directed prevertical setae . PARATRYPHERA Brauer & Bergenstamm 



- Scutellum with four pairs of marginal setae (lateral setae present, strong). Inner 



vertical setae converging and crossing before their apices. Humeral callus with 

 the three main setae standing in a triangle (? Z. pulchra). Pre-alar seta con- 

 spicuously longer and stronger than first post ia seta. ° with a pair of strong 

 outwardly directed prevertical setae ........ 3 



3 Upper occiput with black setulae behind the postocular row. Mid tibia with one 



isolated ad seta. Abdominal Ti + 2 and T3 without median marginal setae. [$ 

 unknown] ........ 'Zenilliana' pulchra Mesnil 



- Upper occiput without black setulae behind the postocular row. Mid tibia normally 



with two or more ad setae (one of the normal two sometimes very reduced or 

 lacking). Abdominal Ti + 2 and T3 with median marginal setae (sometimes 

 reduced in P. vicarium) .... PHOROCEROSOMA Townsend 



Tribe WINTHEMIINI 



This tribe is best represented in the Oriental Region by the nearly cosmopolitan 

 genus Winthemia, but three other genera occur in the area. Baranov (1932c) 



