TACHIX1DAE OF ORIENTAL REGION 141 



This group corresponds very closely to Mesnil's (1952a) subtribe Erythrocerina, 

 but there are certain differences. Elodimyia Mesnil is excluded, as it possesses 

 few of the characteristics of this group and appears undoubtedly to be closely 

 allied to Aneogmena (see A neogmena-gr oup) , and Bactromyiella is excluded because 

 - as emphasized elsewhere (p. 135) - it is probably a blondeliine and not an eryciine; 

 certainly Bactromyiella possesses few if any characters that justify its close associa- 

 tion with Erythrocera. 



The genera Diglussocera and Hapalioloemus are extremely similar to each other 

 and possess all the features typical of the 'Erythrocerines', and are therefore included 

 in this group; they were unknown to Mesnil (1952a) and not placed by him, but it 

 is certain from examination of the type-species (and only species) of Mesnil's (1957) 

 genus Boromyia that this nominal genus is the same as Baranov's Hapalioloemus 

 and Boromyia is accordingly here sunk as a synonym (syn. n.) of the latter. It 

 is uncertain whether Diglossocera and Hapalioloemus should be maintained as 

 separate genera, and I think it likely that the latter will have to be sunk into synonymy 

 with the former when more is known of this rare complex. A remarkable undes- 

 cribed Australian species of the complex has been seen (<$ specimen in ANIC, Can- 

 berra) which conforms to most of the characters of Hapalioloemus (having proclinate 

 orbital setae in the <$, the facial ridges setulose on most of their height and easily 

 visible in profile, and three humeral setae) but which has extraordinarily modified 

 3 antennae and therefore recalls the condition in Diglossocera, although the antennal 

 modification is of a different kind (consisting of many long biramous or multiramous 

 processes arising laterally from each side of the posteroventral edge of the third 

 antennal segment, the whole facial region being enlarged and deeply sunken to 

 accommodate the numerous long branches). The existence of this weird Australian 

 species, as yet unnamed, suggests that there may be other species still to be found 

 whose characters cut across those that define Diglossocera and Hapalioloemus, 

 and that ultimately Diglossocera will require to be widened and redefined to accom- 

 modate them. At present the Australian species alluded to comes closest to 

 Hapalioloemus and could be described in this genus. 



Crosskey (1967c) established that Sigelotroxis Aldrich is a synonym of Atractocerops 

 Townsend, and examination of Frontiniellopsis Townsend for the present work has 

 shown that this name, too, must be sunk as a junior synonym of Atractocerops 

 (syn. n.) Hence the nominal genera Boromyia, Frontiniellopsis and Sigelotroxis 

 all belong in the Erythrocer a-group, but each is a junior synonym. 



Many forms in the Eryt hrocer a-group are distinctive among the Eryciini, and 

 among the Goniinae with large pre-alar seta in general, because of the unusually 

 short and broad wings in which the apical part of the subcostal cell (i.e. the cell 

 between veins Sc and K^ is often very much wider than usual (for example in 

 Elodia and Hapalioloemus from the Oriental area). Other features found in certain 

 members of the group that are unusual among Eryciini include the presence of 

 two pairs of lateral scutellar setae (normally only one pair in other Eryciini and 

 none in Dolichocolon) , the frequent occurrence of fine setulae along the apical part 

 of vein R v and the unusually rotund abdomen. 



