36 RUBIACEAE 
= Galium muricatum Wight. Humboldt Bedstraw. Fig. 5012. 
Gali Wight, seh oi 1900. 
?Galium chartaceum Wight, 1 
Diffuse or a tated ‘giro perennial with at aig stems 3-12 cm. long, the stems 
datieon: sew shining. Leave whorls of 4, sprea py sh than on ee oe or Be aciges 
them in the more tufted thay 5-9 mm. long, firm re, “a rk g shining, n wly to 
broadly elliptic and sometimes obovate, short- aking Fo margins Papa ciliate with aici 
ascending hairs, the upper surface sparsely short-hispid with appressed hairs to glabrous, the lower 
surface glabrous or with occasional hispid hairs on the midvein; inflorescence axillary or terminal 
with slender pedicels, in small clusters of 2 or 3 flowers, little surpassing the leaves; corolla 
th cpboeny white, 1.5-2 mm. wide; fruit glabrous, black in herbarium specimens, fully mature fruit 
esl or ss slopes, Arid Transition Zone; Agness, Curry County, Oregon (Peck, Henderson), an 
Bien and Mendocino Counties, California. Type locali ity: “Westport, Mendocino County.’”’ June-July. Much 
Lisanti the more tufted forms of G. californicum and apparently some forms intergrading with that species. 
23. Galium nuttallii A. Gray. Climbing Bedstraw. Fig. 5013. 
Galium suffruticosum Nutt. in Torr. & Gray, Fl. N. Amer. 2: 21. 1841. Not Hook. & Arn. 1833. 
Galium nuttallit A. Gray, Smiths. Contr. 35: "80. 852 
Dioecious perennial with 1 or more slender stems arising from a woody root, pie 
below and elongate with long internodes, reclining or climbing on shrubs and rocks, 1.5-2 m 
length, coikpacts much branched above and leafy, the seasonal stems ge — ches bertmcceoes, 
more or less retrorse-scaberulous. Leaves in wher s of 4, usually rat becoming thicker 
in age and yp a ran eee: ees ovate: ee ng 2 broadly linear. anal . a rrowly 
linear-oblon n linear in s «fot ms), arely obtuse, the u urface 
= caaeaaiy hispid. “s gittinars he margins inroled a. age Caitty to denaely sabres with 
ually retrorse hairs; staminate inflor n few-flowered cymules in gl hg ie 
sineitate inflorescence solita from jet axils of ‘tea aie wie pedicels 2-4 m ved rolla 
inant i yellow, 2-2.5 mm. broa obes narrowly ovate- Eo aate:. frui pent g 
eter, often 5 1 aescl Gerelonhig pearly white when mature, tert tinged 
with lavender turning black w n dry 
Shrubby hillsides, Sonoran falc vate n and LS ag i Oregon, south through the Inner and 
Outer Coast Ranges to northern Lower California and from Siskiyou and Shasta Counties south along the Sierra 
Nevada foothills. Type locality: San Diego, California. MarcicAce. " [ators yades with G. iolendork are frequently 
found in its gabe limits and along the Sierra Nevada foothills. 
sulare Ferris, Contr. Dudley oe th. . figs. 2a—-c. 1955. Habit as in the 
thick in texture and bearing a cusp. Santa Rosa, Santa fae a d Santa Catalin Islands, California. Type 
cality: Santa Catlina Island, Los Angeles County. 
24. Galium bolanderi A. Gray. Bolander’s Bedstraw. Fig. 5014. 
Galium bolanderi A. Gray, <i Amer. pag a 350. 1868. 
Galium margaricoccum A. Gray, op. cit 1878. 
Galium arcuatum Wiegand, Bull. iba aay Mee 395. 1897. 
Galium trifidum var. Sopliidete um A, Gray, loc. cit., as a synonym. 
Stems loosely branched above, aes to many arising from woody roots, ey angled 
and smooth, more rarely scaberulous, 1.5-4 dm ‘Hay gh, pes oe = — stems somewhat re- 
clining, the new growth with lo ong ‘internodes, stem aching a le ct ah of 7 
Leaves in whorls of 4, sometimes in pairs above, Pe i 4 aie slice, acute at the apex and 
often apiculate, 8-22 mm. long, those of the main stems the longer, thin in texture and homie 
age and dar ied spe r sparse 
5 
ae 
Te 
pedicels 2-7 mm. g, spreading or arcuate; corolla purple = greenish purple, 
the lobes prite Gaeaiice: : pe Fey or pulpy, pearly white when ripe, sometimes hat < one e carpel 
developing, dark in dried spec 
Dry, open, wooded slopes or es ie Transition and Canadian Zones; southern Oregon in the Siskiyou 
Mouktuas region south in oo Coast eal of California to the mountains of Glenn and Lake Counties and 
Mount Hood. Sonoma County, and in the 7 Hexase to Madera County. Type locality: Mono Trail, Yosemite 
National Park. Collected by Bolander. April—Jun 
25. Galium pibens A. Gray. Gray’s Bedstraw. Fig. 5015. 
Galium pubens A. Gray, Proc. Amer. Acad. 7: 350. 1868. 
Galium culbertsontt Cone. Leafléts Bot. Obs. 1: 80. 1904. 
Stems srupscides —<—— from a woody caudex, usually erect but de diffuse ads much branched, 
3-7 dm. high, mo: T less pbs ad with sprea ae & grayish hairs and also scaberulous, rather 
sparsely leafy on pis main stem with internodes 3— ong oO ng much-branc ~~ plants with 
short internodes. Leaves in whorls of 4, firm, obiong-ovate, acite, or linear-oblong an ae on 
much-branched plants, firsts = both surfaces and ous, sometimes obscurely so the 
often inrolled margins, 10-12 mm. long and barely era as long on the ultimate eoucher: 
