62 VALERIANACEAE 
Am shrubs and bushes in shady places (usually with marine influences) at low altitudes, Upper ona 
and Hu =a Transition aa British Columbia, Washington, Oregon, and northern California south along the 
coast to Monter rey and San Luis Obispo Counties. Type locality: ‘‘no orth-west coast of North America,”’ Collected 
congesta subsp. nitida (Heller) Morey, Contr. Dudley Her b. 5: 120. 1959. (Plectritis nitida 
Heller, Muhlenbergia Hy 328. 1907.) Differing from the typical subspecies in the vernicose or mare fruits Mee 
the keel smoothly vas ad - ae the bagi when present appressed and connivent top and bottom; walls of the fruit 
usually more or les e fro he embryo. Slender decumbent plants of more or or Fela wet pan 
poiected 3 the coastal lowland fre Monterey to Fort Bragg, California; rarely elsewhere. Type locality: Crystal 
prings 
Plectritis congesta subsp. ag ea gt (Fisch. 7 as y.) Morey, Contr. Dudley Herb. 5: 119. 19 a 
(Plectritis brachystemon Fisch. & Mey. Sem. Hort. Petrop. 2: 47. 1835; Betckea major Fisch. & 
Apn: Sci, Nat. JI... §: 189. 1836; Plectritis samolifolia Toeck, Bot. Jahrb. < 37. 1882, in part; V alerianella 
anomala A. Gra , Proc. Amer. Acad. ‘S35. V. aphanopter ra A. Gray, cit.: V. magna Greene, Proc. 
Acad. Phila. 1895: 548. 1895; Plectritis iivales "Suksd. Deutsch. Bot. ask 15: 144, Biincd i P. gibbosa 
Suksd. Ws denda 1: 8. 1927.) Differing from the typical subspecies in the smaller flowers (1.0-3.5 mm 
long), wh may lack a spur, and which have a tube 0.5 mm. long or longer, and in the jpcils tubular-funnel- 
form cmos. In brushy hot relatively open, pomp te “habitats scene ag ase asain to Monterey County, Cali- 
fornia. Type locality: ‘‘prope coloniam Ross in California,’? Sonoma Cou 
2. Plectritis macrécera Torr. a art White Plectritis. Fig. 5063. 
Plectritis macrocera Torr. & Gray, : 50. 
ue ra etchleriana Suksd. Detitscti, Bot. Misetix: 15: i aay. 
ligera ostio latata Suksd. loc. cit. 
pb ie jepsontt Suksd. Erythea 6: 24. 98. 
Plectritis arate! Jepson, Fl. W. Mid. Calif. 475. 1901, 
Plectritis collina Heller, Mutter 2! 329.;,1907. 
lants slender, 1.0-6.5 dm. tall. Leaves obovate to ovate-lanceolate, diminishing in size 
upward, pe etiolate below, tag above, entire to coarsely serrate; inflorescence verticillate (some- 
spe 1 i 
aligy ornot at all expanded; fruit pale straw-yellow to red-brown, 2-4 mm. lon ore ess 
pubescent, the hairs stout, clavate to glandular-c ylindric, rarely obuisely donned: oe ‘erga ‘often 
collapsed on drying, or long and curly; keel rounded to rather angular, obtusely angled, often 
a dorsa : 
n na 
with a pair of equatorial pecoeget wings thick, expanded, or entially obsolete, bounded 
with a broad, cylindrical, usually grooved, ok ace thickening, came nt basally and apically 
to an equal degree and prese sora a pete t is broader than tall ; Cavlelchil incumbent, very 
aaa: veined to the third order en expanded 
ed, but not brushy habitats, Upper Sonoran and Humid Transition Zones; San Fra 
cisco 4 Po tase oe Citote from sea Hey o 4,000 feet, with pied pk kee sanoms to southern if cthineton 
and southern California. Type locality: “California.” Collected by Douglas. April—-Ma 
Plectritis macrocera subsp. grayii eam) Morey, Contr. Dudley Herb. Ps 120. 1959. (Aligera grayi 
Suksd. Deutsch. Bot. Monatss. 15: 147. 1897; A. mamillata Suksd. loc. cit.; Plectritis macroptera Rydb. il 
Rocky Mts. 818. 1917, not Suksd.) Differing from the typical subspecies in that the wings when expanded a 
thin and eth a narrow, but usually obvious marginal thickening which is only rarely or ape So igre groove a 
wings are connivent top and bottom to present a large circular orifice; and there is usuall median longitudinal 
ridge on the ventral surface of the fruit which usually bears a multiseriate row of ‘sti - clavate or cylindrical 
bristles. In open, moderately shaded, but not brushy habitats in piedmont and moist, low, montane peacoat 
north-central California northward to the Columbia River and its A ro nslveg eastward to southwestern Montana 
and sth hea to southern California, Nevada, and Utah. Type locality: California. 
3. Plectritis cilidésa (Greene) Jepson. Long-spurred Plectritis. Fig. 5064. 
Valerianella oe it We Proc. Acad. Phila. 1895: 548, 1896, 
Aligera macroptera Suksd. Deutsch. Bot. Monatss. 15: 146. 1897. 
ont oe pany op. cit. 147. Not Torr. & Gray. 
sa Jepson, Man. FI. Pl. — O71... 1925. 
i: si Pst Werdenda 1: 
Plants slender, 1.0-5.5 dm. ta = Leaves obovate to oblong-fusiform, petiolate —— sessile 
above; nae wage to verticillate-spiciform ; pers deep pink, with dark s at the 
base of the ventral lo 5 mm. long, — and narrowly infundibuliform, bilabiat jate, the 
spur slender, usually ey ‘the ovary, the tip not eh ag ; fruit pale straw- 
ellow to brown, 34 mm. long, more or less es the hairs stout ot iff, cylindrical, rome 
or bluntly pointed, rarely somewhat lig eae and not readily ‘collaps ed on drying; 
i than dee 
and stretched from ridge to inner wing-margins, often wit of equatorial mammillae; wings 
variously expanded, rarely obsolescent, bounded with a pa ae dy "ean grooved, marginal 
thickening, connivent tegey # and connivent to more or less spreading above; cotyledons seabed, 
more or less conspicuously veined to the second and third order when expanded. 
In open sunny habitats, grassy hillsides, and stream margins, Upper Sonoran and lower Humid Transition 
Zones; central Cali fornia at higher altitudes in the interior from San Benito County north to Mendocino PSone 
= pons Tuolumne County po to Monterey County; rarely elsewhere. Type locality: Oakville, Napa County. 
pril— 
Plectritis ciliosa sub: nsignis (Suksd.) Morey, Contr, Dudley Herb. 5: 121. 1959. (Aligera insignis 
uksd. Deutsc ie Faia oe i 146. i897) A rubens Suksd. loc. cit.; A. elliformis Suksd. W. Amer. 
Sci. 12: 54. 1901; Plectritis davyana Jepso'! _ W. Mid. Calif. 475. 1901; ligera intermedia Suksd. Wer- 
