120 COMPOSITAE 
2. Helianthella quinquenérvis a agg A. Gray. Nodding Helianthella. 
ig. 5169. 
Helianthus quinquenervis Hook. Lond. Journ. Bot. 6: 2. 1 : 
Fe escusinee quin inquener vis A. Gray, Proc. Amer. Acad. 19: 10. 1883. 
aves ella enervis var. artzonica A. Gray. Syn. Fl. N. Amer. 12: 284. 1884. 
I] la Greene, Leaflets Bot. Obs. 1: 148, 1905, 
Per enni ial we a stout branching caudex, the stems stout, erect, 5-15 dm. ae, Kapaa or 
becoming glabrat a Below. conspicuously but eather sparsely hirsute above. Leaves opposite, 
: a omi i 
up to 50 cm. long, wi 1 
at each end, the basal and lower leaves petiolate, the upper 2 pairs sessile and subclasping ; heads 
long-peduncled, mostly solitary, = acied axillary and short-peduncled, ere: res ie Paes 
ovate-lance prong acuminate, 1. m. long, ciliate-marg ined disk 2.5-4 cm. wide; ra out 
2 ong; recept Sidhe rants thin, soft and sc arious ; achene =e mm. mie ite. 
margined, Sree: -hairy, the broad pappus of 2 slender awns 4-5 mm. line with well-developed 
squam 
Mountain meadows and boggy areas in the forests, Canadian and Huds sonian ares Boange in 
States from the Warner Mountains, Lake Coders. Oregon, and eng cs occurring r elevations in “the 
mountains southeastern Oregon southern Idaho, Montana, and the Black Hills of Seutke Dakota presi to Chi- 
huahua and Nuevo Leon. Type locality: Sooo Natrona County, Wyoming. (See , Am Midl. 
Nat. 48: 20. 1952.) Collected by Geyer. July—Aug 
3. Helianthella californica A. Gray. California Helianthella. Fig. 5170. 
Helianthella californica A. Gray, Pacif. R. Rep. 4: 103. 1857. 
Perennial from a branching caudex; stems several, a ee slender, simple or rarely erect- 
bie: sparsely he with prewar a to spreading hairs, 1.5-6 dm. high. Leaves mostly basal 
or subbasal and opposite, those of stem only 1-2 pairs oe 1-3 alternate ones above; petioles 
er srarrey hae: 3-10 cm. long; blades linear to lanceolate or egg gen 5-20 cm. long, 
acu at each end, entire, feather-veined or Piolinerved near middle, age firm, sparsely 
peianance @ 0B oor ah with subapp =. ae peccading hairs espec cially aioe and on margin, 
becoming glabrate or eeationd lou eads solitary, long- soup tel, 3.5-5.5 cm. yies 
involucre often subtended by 2- 3 Uapoole ee tacts 2 ong, subeq a 1- 
fee the phyllaries lanceolate, 4 poe hirsute-ciliate, more or ey hirsute Morente son or 
uarrose; rays about 12-16, 1-2.5 cm. long; disk yellow, the receptacular ee oa Tpit 
tones at tip; achenes cuneate- ‘obovate, dieale notched at apex, Praag ous; pappus 15. 
Open places on ridges of the Coast rags Upper Sonoran Zone endocin d Glenn Counties, California, 
— ferourh Santa Clara County. Type locality: Napa Valley, ie Camiy. ‘Collected by Bigelow. March- 
wa ianthella californica var. <p octal (Greene) Jepson, Man. Fl, Pl. Calif. 1081. 1925. (Helianthella 
nevadensis Greene, Bull. Calif. Acad. 1: 1885.) Lea ae pid cost puberulent; achenes obovate-oblong, often 
puberulent toward the apex, marginal wine fee: awns 2, slender, ee 1- long and often having 
between them a border of much shorter lacerate squamellae. Mostly i en forests iy the ponderosa pine belt, 
Jackson and Klamath Counties, Oregon, to Modo ¢ and Shasta Counvies: California, a ae Pe so through 
the Sierra Nevada to Tulare pees Kern Counties, and in adjacent Washoe County, Nevada; also in the inner North 
Coast Ranges from Trinity County to Lake County. Type locality: ‘in the higher Sierra,” Coliform. 
Helianthella californica var. shasténsis ~ A. Weber, Amer. Midl. gp 8: oi as e Leaves poe 
appressed-pubescent; achenes Pepin obovate, marginal wings narrow, broader at the a s 2, about 1 mm 
ae Open forests in the Siskiyou and Trinity Mountains, Siskiyou — “Trinity Pouitics Calilorna. ‘ine 
locality: Moa t Eddy, Siskiyou County. Intermediate forms between this variety and the preceding have been 
collected in adi: jacent Shasta County 
4. Helianthella castanea Greene. Diablo Helianthella. Fig. 5171. 
Helianthella castanea Greene, Erythea 1: 127. Pisa 
Helianthella cannonae Eastw. Zoe 5: 82. 1 
Helianthella castanea var. cannonae Jepson, i Fi Pi. Calif: 1081. 192%; 
Perennial, from much-branched caudices, 1.5-4.5 dm. high, similar to H. ave but with . 
larger leaves and more leafy-bracted involucre ; ; stems porch: or ascending, simple or rarely erect- 
branched, 14-45 cm. high, stoutish, hirsute with spreading hairs. re chiefly basa or sub- 
3-8 and alt e ( y wi ir b 
cm la 
obtuse or acute, long-cuneate at base, entire, — at gars b tacierwaieee or usually ‘ipl 
nerved near middle, sparsely hirsute chiefly beneath and on pee of heads cgi long-p 
involucre subtended by about : usually elliptic portend bracts about 4 cm. long, about Bhesinte. 
about 2 cm. high, +; oh Maries yore or lance-oblong, acuminate hispid-ciliate 
and hispid or subglabrous ‘sotealte inflexed over disk after anthesis; s 12-20, cm. 
long; disk yellow; achenes broadly obovate or sedis bicular, with s mall t eoihiad notch, glabrous 
not wing-margined, 8-9 mm. long, 5-6 mm. wide, » distinctly thickened medially when matur 
pappus none or 2 minute teeth, without squamellae * 
Grassy hillsides, Upper Sonoran Zone; Alameda and Contra Costa Counties,  sicpgtlagae San Francisco 
Caz, Congdon. Type locality: summit of Mount Diablo, Contra Costa County. April- —May 
7. ENCELIOPSIS (A. Gray) A. Nels. Bot. Gaz. 47: 432. 1909. 
Scapose xerophytic perennials with stout root and branched caudex, the short branches 
bearing tufts of leaves and monocephalous, essentially naked scapes. Leaves densely 
