240 COMPOSITAE 
the feo > somewhat contorted, not flat, rachis about 1 mm. broad; heads 2 or 3, whitish, 
12-14 mm. high, terminating the leafy stems, the peduncles short; phyllaries cap m. long, 
obtuse or r spatula ate; — lar- fap oe aad sparsely nate: coro olla 
puberulent with occasional glands ; achenes 6-7 mm. long, strigose; peraeae Spalesé 10-46, itacal: 
2-5 mm. Tong sbiGae or linear and rounded at the a i 
en serpentine slopes, Can and Hudso  Tilews ; Wenatchee Mountains in Chelan and Kittitas 
Céiattee: fA oy si osha = locality : omen af Beverly "Creek, Kittitas County. June—July. 
8. Chasers douglasii (Hook.) Hook. & Arn. Hoary Chaenactis. Fig. 5394. 
Hy papp a! ii Hook. Fl. Bor. Amer. 1: 316. 1834. 
pe tis d te) za Bot. Beechey 354. 1840. 
carphus ps [Re tt. s. Amer. Phil. Soc. IT. 7: 
peat os suksdorfit Ae 2 Cane Dudley Herb. 3: 108. — an es Sea 7. 1940. 
igh, stems densely or sparsely tomentose, also 
iennial or perennial herbs 
am ee bas a ee below the 
Stout, b 1.5-6 dm 
ane glandular below the inflorescence, single or few 
biform Ge aifhresciice. Basal eaves 5-1 . long, 2-3-p stem-leaves reduced ad me 
dissected, all thick in texture, m or less euentnee rulandnine: Sal es “alin ate lobes obtus 
con d and so hat Pevolite: the leaf thus not flat; heads large, 50-70-flo , white r 
pinkish; phyllaries 11-17 m g, the outermost shorter, narrowly oblanceolate or oblong, 
granular-gl lar, more or less woolly with soft, slender, jointed hair also me 
coarse straight hairs on the back and margin and also o e peduncle; ee ee hairy on the 
outer surface, the tube glandular ; achenes 6-8 mm. long, densely hirsute; pappus-paleae unequal, 
10- » a near or oblong about — — the length of the corol 
oc andy, sagebrush a , Arid Transition Zone; Yakima County, Washington, south to Jef- 
ferson “County S Ordena Type loeality: se Celilo Falls on the Columbia River. June—July. Douglas” Pin- 
cushio 
at axon which is part of an extremely variable and widespread complex which p tege from the avail- 
ric seach data, is quite pees ” Beers conditions; consequently ecological as well as regional forms 
o be found. In eastern Was eastern Origen plants occur that are intermediate in their character- 
pps betwee C. douglasit var. git Gad C. douglasti var. “sialaetlonh a These forms that cygnetatars might 
be considered od be C. douglasii var. achilleaefolia have more flow ead and some coarse ight hairs 
on the phyling ie: 
All of the taxa of the douglasti complex have to a greater or less degree a adi ah ory 2 leaf-character. 
The leaves do not — a plane Ps ae but rather a contorted one, due partly to the orientation of the 
nn nd a tendency to become revolute. This is specially noticeable in the voloced forms pe higher eleva- 
tions. 
Aig enactis douglasii var. achilleaefélia (Hook. & Arn.) 
Roc ts. 577. 1909, (Chacnactis appceci ole Hook. & Arn. Bot. Beechey ah 
leafolio Nutt. Trans. Am Phil. Soc 376. 1841; Chaenactis imbric ‘a Greene, 
1912; C. bracteata Gs ene, op. cit. Tine? é. cheilanthoides Cresits op. 7 . 
t m . i 
stud ab co 
A. Nels in Coult. & Nels. New Man. Bot 
1840; Macrocarphus achil. 
Leaflets Bot. Obs. 
h; 
5 te landular-puberulent as well loosely canescent; corollas, F tanes s, and 
pappus- pale ae as in C. douglasti douglasit, but the pappus- py especially of plants of the southern ex- 
pies Ot the range, usually sechatk - Bs 20g the length of the nggriges Plains and dry slo opes, British Columbia 
o Montana and south to northern Ari and southward east of the Caeade Decarhaing in Washington and 
Sane to northeastern acs rnia; on a Sierra Nevada principally on the eastern face and the desert ranges 
as far south as Inyo Count 
As here identified ae penilleasiete is the mon segregate of the douglasii complex and m 
uded under the Also peiecane hy with named ee from nigher elevations as oe 
riants are incl 
a rebase of lower <a = ae and rolaies are frequently found. Type locality: “Dry plains of the Snake 
Country.” Collected by Tolm 
Chaenactis Sangtonis var.  aplantalbce Cronquist, vor. — Pacif. Northw. 5: : 5. Annual or 
biennial ©) about 3 dm. or ace high, differing from the two preceding taxa in green er foliage, stipitate- 
ndular odanal and nd consistently pink * ah nu of flowers in each head as in 
regon, and adjacent 
pedun a s 
gi ‘dong lasii var, achilleaefolia. seek canyons in oS Washington, idee pte ak 
Washington. Dusty Mai 
Chaenactis ii ane var. monta Eee E. Jones, Proc. Calif. Acad. II. 5: 700. 1895. (Chaenactis pana- 
mintensis Stockwell, Sasa Bags pe He oe Seeks ass ee comer var. nana Stockwell, ae cit. 117; C. doug- 
—_ var. gem yg tthe , Madrofio 7: “St 1943.) ennial, leafy-stemmed plan 
th 1 dm. ee SN = oa ror reddish, erect, aati: freely branching, 
om a eabtercane branched caudex, the be vie tons thick 
bess io great thickne s i. S aikate of tates slopes); basal and lower leaves many, tending to eh ps 7a, 
r leaves reduced in oe ze; a ga Hesse: sely branched as ob a 
ee shyla deb unequal, 8-10 mm. long, granular 
oftén also woolly- a espec ily Tucked a pene _pappus-pa aleae of the in 
3 ee Pa 
App ont only an ; cnainn at higher Shey ations of a achilleaefolia which has developed the ch 
of shart gover and oe. grow “vege Be haenactis pedicularia ‘Greene sé ony 4: 98. 1899), C. “ongustfolia 
Greene (Leaflets Obs. 223. 2); °C — Pg (N. Amer. Fl. 34: 72. sit be 0), and C. 
Stockwell (Contr. spualey “a a at eet s obably synonyms pe this su balpine — 
t of the le afy branching Ppa 5 wih ce vident inter- 
Peak, Panamint Mountains, Califo d, -—_ din . one- ehalf on ee “the. crn Nevad of the corolla. pn tan 
- 1 rni 1a, an ace ing t ‘oc t' 
Olancha Peak, Tulare County, California (Munz 15287), appeers to be this form. en 
Chaenactis Jowdiaati var. rubricailis (Rydb.) Ferris, Poca fete ei 5: ae Menge rtd 
rubricaulis Rydb. N. Amer. Fl. 34: 72. 1914.) Perennial 1.5—3 stems red. few (1-6), 
nearly glabrous with traces of a lan ate tomentum especially any he Pies of the Pe sei slandular on 24 
t . long, the 
pec 
—-- - slender root often hori rizontal, the stems Shaecae Leaves mostly basal, abou 
aves . So ipi i 
ucres 
hecomin; glab : ste 
long. rather narrowly linear, obtuse at a usually densely: anduiar- uberulent; ce 
reddish-tipped: pappus of 10 nearly equal, Cohlanceokate paleae. som sl si se, miore th 
corolla. In the upper Arid T n Zones in the sg pied Nesoae from Hone County to Calaveras 
