420 COMPOSITAE 
Arnica chamissonis subsp. foliosa var. ereanenn (Greene) Maguire, Amer. Midl. Nat. 37: 140. 
ome bernardina bag Pittonia 4: 170. a es var. bernardina Jepson, Man. Fl. Pl. Calif, ins, 
es — nar: Mountai ains, California. Shy ge ocality: Bear Valley. 
chamissonis << foliosa var. incana Gray) re Fl. Alaska & Yukon a A 1950. 
Pel4s-1 re 
a Greene Cc 
the s end of th 
i oe ag Beiutains of St thern Wa line gton, ex xtending eastward occasionally fate pets so Utah and south- 
western Idaho and apparently sometimes northward to southern Yukon. Type locality: Sierra Valley, California. 
Arnica chamissonis subsp. foliosa var. foliosa. Range of the subspecies except the mountains of southern 
aiicons. 
pet hosaadinggan ie m of A. chamissonis subsp. foliosa, which approaches As parryi var. i in its some- 
me Ror iavonticral bracts ape, in its more or less reduced cauline leaves, has been = 
pene a he taaii of a few collections Lhe the Sierra Nevada as var. jepsoniana Marcie, Aare. Midl. Nat 
37: 140. 1947. The real biological nature of these s sche iid remains to be determined. 
2. Arnica longifolia subsp. aon ead Maguire. 
Seep- 
as ee Fig. 5 
Piper, Proc. Biol. Soc 06. 1920. 
Arnica louse ibe weirichei anges eda 4: i 1943. 
Plants densely tufted, often in very large clones, with many smaller, re leafy stems in 
addition to the floriferous one s, these commonly 3-6 dm. tall, leafy, the rhiz commonly short- 
ned into a branching caudex. These mostly 5-7 pairs, one or more of the ss pairs connate- 
sheathing, with reduced blade, the others sessile or shortly connate-petiolate, not much reduced 
upward, narrowly lanceolate or lance-elliptic, pal tapering to the acute or acuminate tip, 
or tly 5 a 2 ide; no we 
entire or sometimes slightly toothed, mos 4 -12 cm. long and 1-2 cm. wid well-developed 
basal leaves produced, either on stem or on separa sie short shoots ; herbage more or less scabrid- 
puberulent, at least upward, and usually a “ttle viscid ; heads setaral to rather numerous, cam- 
panulate, sometimes rather narrowly so; involucre mostly 7-10 mm. high, the bracts sharply acute 
or acuminate, glandular-puberulent and generally with a few intermingled, longer, conspicuously 
septate hairs, especially below ; rays mostly 8-13, 1-2 cm. long; achenes subglabrous or glandular 
and hairy; ore or less tawny, barely subplu umose 
Il-drained soil (or rocks : —_ oes Peg: pectag? are salons oist cliffs and river banks, at modera 
o hig 
1 ether 
and oe ‘Alber ay gm peaitee: Mount Rainer National Park, Washington. July— ee 
58 seit amplexicaulis Nutt. Streambank Arnica. Fig. 5721. 
Arni. bl. lis Nutt. Trans. Amer. — Soc. IT. 7: 408. 1841. 
ica macounii Greene, hing 160. 190 
Arniea abe oakte Sal Rydb. Mem. N.Y. Bot. re 1: 434. 1900. 
eene, Ottawa Nat. 15 or 1902 
ria ciliate Rydb. N. ene Fl. 34: 351. 1927. 
Arnica macounii var. aspera G. N. Jones, Oa Wash. Publ. Biol. 5: 256. 1936. 
Perennial from sittick coarse, freely ing rhizomes, mostly 3-8 dm. tall, more or less glandu- 
lar and hairy, especially upward, or s ate ial beds leafy. Le eaves all caine mostly 5-12 pairs, the 
lowermost often reduc oe e other: ers elf developed and generally no h redu rd, nar- 
rowly lance-elliptic to ecg € or ovate, sania or the lowe seth hota: etiolate, more or less 
toothed, only cm. long and 1-3.5(4) cm. — heads generally several, campanulate ; 
involucre mostly 9-15 mm. high, its bracts sharply acute or acuminate; rays 8-14, pale yellow, 
1-2 ~ long; achenes spaniels hirsuite and sometimes  Yeadisla ar; pappus tawny, subplumose. 
am banks moist woods, Canadian and Transition Zones; Aiaake to western Montana, northeastern 
sl — to the southern Sierra Nevada. Type in locality: “On the nie oof the Wahlamet, at the 
* me 
i keke var. piperi St. er & Warren in St. John, Proc. Biol. Soc. Ph a 36. 1931. 
(A, Rrtica Amer. Fl. 34: 349. 1927.) Leaves very broad, up to 6 or reput edly 8 wide; in- 
volucre a genes hairy cad less glandular; otherwise much as in typical A. Seniesa but ‘ess variable. 
Columbia River Gorge in Oregon and Washi ington i Type "Pocatiog Cape Horn, Washingt 
4. Arnica méllis Hook. Cordilleran Arnica. Fig. 5722. 
Arece mollis Hook. Fl. Bor. Amer. 1: 331, 1834. 
: e, Pittonia 4: 36. 1899, 
rnica rt levis Claene. ~ cit. 163. 1900. 
— calle teak Greene, op. cit. 165. 
Arnica mollis var. keahicvina Smiley, Univ. Calif. Pub, Bot. 9: 386. 1921, 
rennial from freely rooting rhizomes, which ma ortened into a loosely branched 
caudex, 2-6 dm. tall, variously puberulent to long-hairy Bp tag Cauline Lene 
poe ee, a lower commonly the largest, all sessile, or the lower often s petiolate, variously 
elliptic, or lanceolate to oblanceolate or bovate, irregularly denticulate (or nl 
a "Vita loped, petiolate, basal leaves often enh i Sg © few or se FS gorge fg 
ca ulate or broader, me rs times as s e; involuc ‘Doe 
places at aidan to hie cage in oe mountains, Canadian and Hud ian Zones; Alberta and 
British Combi to southern Colorado, southern Utah, and the sout ae faa of the: Sie erra Nevada in Cali- 
e higher mountains of. Washington and Oregon but absent from the California Coast Ranges, 
