NEW SPECIES OF CROCIDURA 



105 



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Fig. 2 Ventral view of cranium from left to right of C. attenuate! BM(NH) 191 1.9.8.26, C. hiltiana BM(NH) 1994. 113 and C. fuliginosa 

 BM(NH)1933.4.1.183. 



range of C. fuliginosa (see Table 1). It is readily distinguished 

 from both species by its robust, angular cranium, in which the 

 maxillary region is broad, the interorbital region narrow and 

 the anterior part of the braincase markedly angular (see Figs 

 1-3). In contrast, both C. attenuate! and C. fuliginosa have a 

 proportionally narrower maxillary region, broader interor- 

 bital region increasing noticeably from anterior to posterior 

 and a more rounded braincase that is evidently broader than 

 the maxillary region. Lambdoid crests are more or less well 

 developed in both C. hilliana and C. fuliginosa, but they meet 

 at an acute angle at the midline in C. hilliana and a shallower 

 angle in C. fuliginosa; lambdoid crests are less developed in 

 C. attenuata and meet at a shallow angle. Squamosal crests 

 are absent or ill defined in C. attenuata, poorly to moderately 

 defined in C. fuliginosa but well-marked in C. hilliana. The 

 mandible of C. hilliana is considerably more robust than that 

 of either of the other two species (see Fig. 4). The horizontal 

 ramus of the mandible of C. attenuata is more slender than 

 that of C. hilliana, with a sinuous ventral border; the coro- 

 noid process is considerably narrower and shallower; the 

 ascending ramus is higher and the condyle is higher than 

 broad (ratio of condyle width to height 75.0-93.3). The 

 horizontal ramus of the mandible of C. fuliginosa is longer yet 

 shallower than that of C. hilliana, with a narrower, less robust 

 coronoid process and, as in C. attenuata the condyle is higher 

 than broad (ratio of condyle width to height 77.8-93.3). The 



mental foramen lies below the posterior part of P 4 in C. 

 attenuata and C. fuliginosa but below the anterior of M, in C. 

 hilliana. Dentally the most obvious differences between C. 

 hilliana and the other two species is the comparatively large 

 anterior dentition (I 1 , Un 1 and I,) relative to the rest of the 

 teeth, in combination with the narrow M 3 and the reduced M 3 

 of C. hilliana, differing considerably from the condition in 

 either C. attenuata or C. fuliginosa (see Table 1 and Figs 2-6). 

 In detail the dentition of C. attenuata differs in the follow- 

 ing aspects from that of C. hilliana: I 1 is slender and orth- 

 odont, the posterolingual cingulum is narrow, Un 1 is 

 moderate in size and the distance between the two first upper 

 unicuspids is greater than the breadth of Un', Un 2 is smaller 

 than Un 1 and Un 3 , and the unicuspids overlap only slightly so 

 that the rostrum is moderately long in appearance; the 

 parastyle of P 4 is moderately well developed. M 3 is variable in 

 different populations of C. attenuata; it is medium sized in 

 Indian and Burmese populations and thus readily distin- 

 guished from C. hilliana, and although only narrower on 

 average in the Chinese populations of C. attenuata, neverthe- 

 less, the lingual basin is less compressed than in C. hilliana. 

 The first lower incisor of C. attenuata is moderately slender, 

 straighter and more procumbent than that of C. hilliana; the 

 anterolingual ridge extends for two thirds the length of the 

 tooth and is subparallel to the ventral border of the tooth; the 

 posterior border of I, lies below the posterior part of I 2 ; less 



