108 



P.D. JENKINS AND A.L. SMITH 



Fig. 4 Lateral view of mandible from top of C. attenuata 

 BM(NH)1911.9.8.26, C. hilliana BM(NH)1994.90 and C. 

 fuliginosa BM(NH)1933.4.1.183. 



of C. attenuata either amongst the owl pellet remains from 

 Loei Province or from remains found at M. lyra roosts in 

 Thung Yai-Huai Kha Khaeng. It is therefore uncertain if C. 

 hilliana is sympatric with either C. fuliginosa or C. attenuata. 



Crocidura hilliana does, however, occur sympatrically with 

 a smaller species of Crocidura which proved difficult to 

 determine from the fragmentary skulls in the owl pellets. 

 Allen & Coolidge (1940) collected C. vorax (currently 

 grouped with C. pullata Miller, 1911 from the Himalayas, see 

 Hutterer, 1993) from northwestern Thailand, while a speci- 

 men from Lat Bua Kao, mainland Thailand, attributed to C. 

 fuliginosa by Kloss (1919) is also an example of C. p. vorax. 

 Several skulls attributable to this species were found in the 

 owl pellets from Loei, while a good series was recovered from 

 the M. lyra prey remains from Thung Yai-Huai Kha Khaeng, 

 where an additional skull was found in the faeces of a large 

 carnivore. The only other species of Crocidura listed by 

 Lekagul & McNeely (1977) from mainland Thailand was C. 

 horsfieldii indochinensis from Chiang Mai and Khao Yai 

 National Park . Most recently, Davison (1984), recorded C. 

 monticola from penisular Thailand. Neither of the last two 

 species mentioned above were identified from either area, 

 although pellets from Loei Province contained another shrew 

 Suncus etruscus (Savi, 1822), plus a variety of rodent and bat 

 species. 



Since there has been so little systematic collection in 

 Thailand, it is impossible to make categoric statements about 

 the new species, however it seems likely that it is relatively 

 localised in its distribution. Even in areas where collecting 



Fig. 5 Lateral view of left anterior dentition. Left: upper toothrow 

 (I 1 to P 4 ); right: lower toothrow (I, to P 4 ). Top: C. attenuata 

 BM(NH) 191 1.9.8.26; middle: C. hilliana BM(NH) 1994. 119 upper 

 toothrow, BM(NH)1994.118 lower toothrow; bottom: C. 

 fuliginosa BM(NH) 1933.4. 1.178. Scale 1 mm. 



Fig. 6 Occlusal view of left upper toothrow from left to right of C. 

 attenuata BM(NH)1911.9.8.26, C. hilliana BM(NH)1994.121 and 

 C. fuliginosa BM(NH)1933.4.1.178. Scale 1 mm. 



