118 



N. CUMBERLIDGE 



(1991), and can be distinguished from S. flowed by examina- 

 tion of gonopod 1: the raised lobe on the cephalic part of the 

 terminal segment in S. kagoroensis is lower than that in S. 

 floweri, and the outer margin of the subterminal segment of 

 gonopod 1 is slim, while that of 5. floweri is conspicuously 

 broadened (Fig. 2 d, e). Furthermore, the carapace of S. 

 kagoroensis is significantly (P <0.001) flatter (CH/CL = 

 0.44) than that of S. floweri (CH/CL = 0.61). 



Sudanonautes granulatus was redescribed by Cumberlidge 

 (1993a) and can be distinguished from S. floweri as follows. 

 The carapace of S. granulatus is significantly (P <0.001) 

 flatter (CH/CL S. granulatus = 0.51, S. floweri = 0.61), and 

 less widened (CW/CL S. granulatus = 1.41, S. floweri = 1.51) 

 than that of S. floweri (Table 1). In addition, the dactylus of 

 the major cheliped of the adult male of S. granulatus is 

 dramatically arched, while that of S. floweri is only moder- 

 ately arched; the major cheliped of adult male S. granulatus is 

 as long as, or longer, than the carapace width (Cumberlidge, 

 1993a), whereas that of S. floweri is shorter (Fig 1 f) than the 

 carapace width (Fig. 1 a,b). 



Three other species, S. aubryi, S. africanus, and S. ortho- 

 stylis, differ from S. floweri in that the terminal segments of 

 gonopod 1 of these species lack both a raised cephalic lobe, 

 and a distinct longitudinal groove in the caudal view. These 

 three taxa can be further distinguished from S. floweri as 

 follows. The carapace of S. aubryi is significantly (P <0.001) 

 flatter (CH/CL S. aubryi = 0.52, S. floweri = 0.61), and less 

 wide (CW/CL S. aubryi = 1.37, S. floweri = 1.51) than that 

 of S. floweri (Table 1). In addition, the carapace and post- 

 frontal crest of 5. aubryi are a green-brown colour, whereas 

 these parts of S. floweri are uniformly red-brown with a 

 contrasting yellow postfrontal crest. 



The terminal segment of gonopod 1 of S. africanus is thin 

 and needle-like, while that of S. floweri (Fig. 2 d) is wider and 

 has a distinct groove in the caudal view. The carapace of S. 

 africanus is significantly (P <0.001) flatter (CH/CL S. africa- 

 nus = 0.43, S. floweri = 0.61) and less wide (CW/CL S. 

 africanus = 1.38, S. floweri = 1.51) than that of 5. floweri 

 (Table 1). The carapace of 5. africanus has patches of raised 

 warts, while that of 5. aubryi is completely smooth. Finally, 

 the pollex of the propodus of the major cheliped of S. 

 africanus has a large and conspicuously flattened tooth, which 

 is lacking in adult S. floweri. 



Sudanonautes orthostylis was redescribed by Cumberlidge 

 (1993b), and can be distinguished from 5. floweri as follows. 

 The terminal segment of gonopod 1 of S. orthostylis is 

 straight, lacks a visible groove, and curves outwards sharply 

 only at the tip, while that of S. floweri bears a longitudinal 

 groove and curves from the mid point (Fig. 2 d). The 

 carapace of 5. orthostylis is significantly (P <0.001) flatter 

 (CH/CL S. orthostylis = 0.51, S. floweri = 0.61), and less 

 wide (CW/CL S. orthostylis = 1.44, S. floweri = 1.51) than 

 that of S. floweri (Table 1). The frontal margin of S. 

 orthostylis is significantly (P <0.001) wider than that of 5. 

 floweri (FW/CL S. orthostylis = 0.46, S. floweri = 0.38, Table 

 1). The dactylus of the major cheliped of S. orthostylis is 

 broad and flat, while that of S. floweri is narrow. Finally, 5. 

 orthostylis is a much smaller species, maturing at CW 22 mm, 

 compared to maturity between CW 33-42 mm in S. floweri. 



Acknowledgements. I am very grateful to Mr. Paul Clark and Ms. 

 Miranda Lowe (NHM, London), Drs. D. Guinot and J. Forest 

 (MNHN, Paris), and Mr. Trefor Williams (University of Liverpool, 



UK) for loaning the specimens used in this work. The following 

 people are thanked for hosting visits: Dr. H. Feinberg, AMNH, New 

 York, USA; Ms. Ardis Baker Johnston, MCZ, Harvard, MA, USA; 

 Mr. Paul Clark, NHM, London, UK; Drs. L. Holthuis and C. 

 Fransen, NNH, Leiden, The Netherlands; Dr. R. Joque, RCM, 

 Tervuren, Belgium; Dr. R. Manning, USNM, Washington DC; Drs. 

 H.-G. Andres and G. Hartmann, ZIM, Hamburg, Germany; Dr. H. 

 Gruner, ZMB, Berlin, Germany; and the staff of the FMC, Chicago, 

 USA. I especially thank artist Mr. Jon C. Bedick of Northern 

 Michigan University, USA, for all of the illustrations used in this 

 paper. Part of this work was supported by a Faculty Grant from 

 NMU, Marquette, MI, USA. 



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