10 
P.D. JENKINS AND M.F. ROBINSON 
Table 2 Data matrix from Frost et al. (1991: Appendix 2) with the addition of characters scored for two additional taxa, Hylomys megalotis and Hylomys 
parvus 
Character 
Hypothetical ancestor 
Echinosorex gymnura 
Podogymnura aureospinula 
Podogymnura truei 
Hylomys sinensis 
Hylomys suillus 
Hylomys hainanensis 
Hylomys megalotis 
Hylomys parvus 
Hemiechinus aethiopicus 
Hemiechinus hypomelas 
Hemiechinus micropus 
Hemiechinus auritus 
Hemiechinus collaris 
Mesechinus dauuricus 
Erinaceus amurensis 
Erinaceus concolor 
Erinaceus europaeus 
Atelerix frontalis 
Atelerix algirus 
Atelerix albiventris 
Atelerix sclateri 
Tenrecoids 
Soricoids 
Lepticidae (fossil) 
111111111122222222223333333333444444444455555555556666666666777777777788 
123456789012345678901234567890123456789012345678901234567890123456789012345678901 
000000000?000?00000?000000000000000000000000000000000?7000?00?7000000007?77000000000 
000010112211010000020010000000001000100000010000000100010000111001101111110000001 
00001001121101000002010000000000110010001001000100010111010011100110?711110000001 
100101010211100100120100000000001100100010000100020011111111111001101111110000000 
000101010211100100120000000000001100100010000000010000011111111001101111110000000 
000101010211100100120000000000001100100010000000010000111111111001107?11110000000 
100010012211000010120100000000001100100010000000010000000000111001101011110000000 
111201102131011011011001104001110011111101101011120001112111121110210000001130200 
111201102121011011011001103001110011111101101011120001112010121110210000001130200 
111201102121011011011001104001110011111101101011120001112111121110210000001130200 
111201102111011011011001102001110000111101101011120001112010121110210000001130300 
111201102111011011011001102001110000111101101011120001112010121110210000001130300 
110201102110011011011001101101011000111101101011120001112010121110210000000120310 
110201102111011011011001101011010000111101101011221001112010121110210000000011110 
110201102111011011011001101011010000111101101011221001112010121110210000000010110 
11020110211101101101100110101101000011110110101122100111201012111021000000001?110 
110201102111011011011101111011010000111101101011120001102010121110210000000111110 
110201102111011011011101111011010000111101101011120001102011121110210000000110110 
110201102111011011011101111011010000111101101011120001102111121110210000000210110 
110201102111011011011101111011010000111101101011120001102111121110210000000110110 
00000000030001000000000000000000000000000000000000?001000000000000000000000000000 
00000000030001000002000000000000000000000000000000?001000100000000000000000000000 
000000000000000000000000000000000000000000000000000000000000000000000???000?70??? 











Table 3. Selected measurements of Hylomys in millimetres to show variation in size and proportions. The mean, standard deviation and range are 
provided, with sample size in parentheses. 
Character 
Head and body length 
Tail length 
Ratio of tail length to head and body length 
Hindfoot length 
Ear length 
Greatest skull length 
Condyloincisive length 
Upper toothrow length 
Length from I1 to anterior of P4 
Ratio of 11—P4 length to upper toothrow length 
Rostral breadth 
Rostral length 
Ratio of rostral length to rostral breadth 
Braincase breadth 
Braincase height 
Ratio of braincase height to braincase breadth 
' Measurements from Shaw & Wong (1959). 

H. parvus H. sinensis H. suillus H. hainanensis' H. megalotis 
107.67 + 4.50 115.75 + 4.83 1325 5.92 136.71 + 8.28 124.23 + 7.68 
104-114 (3) 111-124 (6) 120-139 (10) 120-147 (7) 115.6-134.7 (4) 
24.00 + 0.82 68.5 + 3.70 2B OOS 09 39.67 + 2.13 86.33 + 3.29 
23-25 (3) 63-73 (6) 19-30 (8) 36-43 (6) 82.8-91.3 (4) 
0.22 + 0.01 0.57 + 0.05 0.17 + 0.35 0.29 + 0.13 0.70 + 0.04 
0.21-0.24 (3) 0.51—0.63 (6) 0.14—0.24 (8) 0.27-0.31 (6) 0.65-0.74 (4) 
23.17 + 0.24 25.88 + 1.02 PIES OES 25.14 + 1.62 20.83 + 0.38 
23-23.5 (3) 24-27 (6) 20.5—23 (9) 24-29 (7) 20.4—21.3 (4) 
18.38 + 0.45 17.71 + 1.41 WSS AAS 22.43 + 0.90 
NIE VOD) 18-19 (6) 15-20 (7) 16-19 (6) 20.9-23.2 (4) 
32.38 + 0.82 34.45 + 0.80 34.65 + 0.60 37.76 + 1.01 
30.55 (1) 31.07-33.64 (10) 33.48-35.61 (7) 33.7-35.3 (4) 36.36-39.23 (4) 
30.98 + 0.82 33/3 = O67; 36.96 + 1.04 
29.09 (1) 30.05-32.36 (11) 32.88-35.01 (7) 35.66-38.56 (4) 
15.21 + 0.42 NG Pas O37) 17.75 + 0.64 17.26 + 0.16 19.89 + 0.47 
14.64-15.61(3) 15.64-16.82 (11) —16.70-18.75 (13) ——-17.0-17.50 (5) 19.44—20.64 (4) 
8.19 + 0.47 8.94 + 0.44 9.79 + 0.46 11.65 + 0.33 
7.54-8.61 (3) 8.15—9.54 (11) 9.05—10.34 (13) 11.40-12.21 (4) 
0.54 + 0.02 0.56 + 0.02 0.55 + 0.01 0.59 + 0.01 
0.52-0.56 (3) 0.51-0.58 (11) 0.54-0.58 (13) 0.57-0.59 (4) 
Slt = (0), 17/ 5.92 + 0.18 6.02 + 0.21 5729 2 0als 
4.98—-5.35 (3) 5.59-6.20 (11) 5.57-6.44 (13) 5.06-5.38 (4) 
9.85 + 0.49 9.67 + 0.29 10.35 + 0.41 13.57 + 0.43 
9.33-10.5 (3) 9.33-10.37 (11) 9.57-10.85 (13) 13.30-14.32 (4) 
0.52 + 0.03 0.61 + 0.02 0.59 + 0.02 0.39 + 0.01 
0.47-0.55 (3) 0.57-0.65 (11) 0.56—0.63 (10) 0.37-0.41 (4) 
13.89 + 0.31 15.60 + 0.86 14.70 + 0.54 14.61 + 0.34 
13.29 (1) 13.42-14.45 (11) = 12.97-15.69 (10) ~—-13.9-15.4 (4) 14.16—-15.06 (4) 
9.06 + 0.34 9.47 + 0.33 8.75 + 0.19 
8.63 (1) 8.55—9.67 (11) 9.33-9.85 (7) 8.49-8.95 (4) 
0.65 + 0.03 0.68 + 0.02 0.60 + 0.02 
0.65 (1) 0.61—-0.70 (11) 0.63—0.70 (7) 0.58-0.62 (4) 
