
The Bulletin of The Natural History Museum (formerly: Bulletin of the British Museum 
(Natural History) ), instituted in 1949, is issued in four scientific series, Botany, 
Entomology, Geology (incorporating Mineralogy) and Zoology. 
The Zoology Series is edited in the Museum’s Department of Zoology 
Keeper of Zoology Prof P.S. Rainbow 
Editor of Bulletin: Dr B.T. Clarke 

Papers in the Bulletin are primarily the results of research carried out on the unique and ever-growing collections of the 
Museum, both by the scientific staff and by specialists from elsewhere who make use of the Museum’s resources. Many of the 
papers are works of reference that will remain indispensable for years to come. All papers submitted for publication are 
subjected to external peer review before acceptance. 
SUBSCRIPTIONS Bulletin of the Natural History Museum, Zoology Series (ISSN 0968-0470) is published twice a year 
(one volume per annum) in June and November. Volume 68 will appear in 2002. The 2002 subscription price (excluding VAT) 
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only price available to institutional subscribers is £79.00 (US $140.00 in USA, Canada and Mexico). 
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Copyright © 2002 The Natural History Museum 
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World list abbreviation: Bull. nat. Hist. Mus. Lond. (Zool.) 
ISSN 0968-0470 
The Natural History Museum Zoology Series 
Cromwell Road Vol. 68, No. 1, pp. 1-50 
London SW7 5BD Issued 27 June 2002 
Typeset by Ann Buchan (Typesetters), Middlesex 
Printed in Great Britain by Henry Ling Ltd., at the Dorset Press, Dorchester, Dorset 
