36 



p. a. Mclaughlin and p.f. clark 



1mm 



Fig. 3 Shield and cephalic appendages, a. Diogenes goniochirus Forest, 

 1956, 9 SL = 2.15 mm, UMZC Nov. 30, 1899; b. Diogenes avarus 

 Heller, 1865,o* SL= 1.53 mm, UMZC Nov. 30, 1899. 



dorsolateral distal angle weakly produced and terminating in small 

 spine, dorsomesial distal angle with small spine, laterodistal margin 

 usually with 2 small spines, and 1 or 2 prominent slender spines 

 ventrally. First segment with 1 small spine on lateral margin ventrally. 

 Antennal acicle not reaching to distal apex of fourth peduncular 

 segment, terminating in bifid spine, with 2 widely-spaced spines on 

 mesial margin. Antennal flagella moderately short, approximately 

 as long as ambulatory legs; each article with pair of long pinnate 

 (pappose) setae. 



Maxillule with endopod lacking external lobe. First maxilliped 

 (Fig. lc) lacking flagellum; endopod fused to exopod. Third 

 maxilliped with 2 strong spines on basis, ischium without crista 

 dentata but with 1 very strong curved spine; merus with 1 or 2 spines 

 on ventral margin. 



Left cheliped (Figs 2a, 3a) with dactyl approximately 0.33 longer 

 than upper margin of palm; cutting edge with row of calcareous 

 teeth; terminating in small calcareous claw, overlapped by fixed 

 finger; outer surface flattened, with short row of 4 or 5 blunt to 

 extremely slender and acute spines near upper margin and 2 or 3 

 spinules centrally; upper margin with row of strong, subacute to 

 extremely acute spines, decreasing in size distally and sparsely 

 interspersed with long setae; inner surface with few widely scattered 

 long setae. Fixed finger with outer surface flattened, armed with few 

 scattered small blunt or sharp spines; lower margin armed with row 

 of strong, blunt or acute spines, sparsely interspersed with long 

 setae, and forming weak curve with lower, similarly armed margin 

 of palm; cutting edge with row of calcareous teeth; terminating in 

 prominent calcareous claw. Palm with outer surface convex, armed 

 with scattered blunt or acute spines, with irregular row of stronger 

 spines beginning near midpoint of proximal margin, curving up- 

 ward, and continuing to near distal articulation with fixed finger; 

 upper margin with 5 or 6 strong subacute or very acute curved 

 spines; inner surface with few scattered tufts of short setae. Carpus 



approximately equal to or very slightly longer than palm; upper 

 margin with row of 4 or 5 strong spines, outer face convex and with 

 slightly oblique row of 3 widely-spaced spines; inner surface gla- 

 brous. Merus triangular; dorsal margin with row of spinules and tufts 

 of setae, 1 much stronger spine at dorsodistal margin; ventromesial 

 with 3 or 4 small spines in proximal half; ventrolateral margin with 

 3 or 4 spinules in distal half. Ischium unarmed. 



Right cheliped (Fig. 2b) appreciably smaller than left. Dactyl 

 approximately equal to length of palm; upper margin not distinctly 

 delimited, outer surface with few spinules partially obscured by long 

 setae; cutting edge with row of fine corneous teeth, terminating in 

 small corneous claw and overlapped by fixed finger. Palm with row 

 of 3 spines and long setae on or adjacent to upper margin, convex 

 outer face with varying amounts of long setae, usually 1 additional 

 small spine distally near upper margin and second small spine near 

 base of fixed finger, lower margin unarmed; fixed finger with row of 

 very fine spinules in midline; cutting edge with row of corneous 

 teeth, terminating in small corneous claw; inner surfaces of dactyl, 

 fixed finger and palm with long setae, most numerous on dactyl and 

 fixed finger. Carpus with long setae and 3 strong spines on upper 

 margin, outer surface convex, with few long setae and 1 prominent 

 spine at mid-distal margin; inner and lower surfaces with scattered 

 setae. Merus triangular; dorsal margin with 1 or 2 spinules and long 

 setae, 1 more prominent spine at dorsodistal margin; ventromesial 

 margin with 2 or 3 small spines in distal half, ventrolateral margin 

 with 1-3 spinules distally. Ischium unarmed. 



Ambulatory legs (Figs 2c, d) with dactyls approximately equaling 

 length of propodi, both dactyls and propodi of second appreciably 

 longer than third, unarmed but with numerous long setae. Carpi with 

 dorsodistal spine and 1 spine on dorsal surface in proximal half 

 (second), unarmed or with tiny proximal spinule (third), dorsal and 

 ventral surfaces with long setae. Meri with long setae on dorsal and 

 ventral surfaces, second with 2 widely-spaced spines, third unarmed. 

 Ischia unarmed, but with long dorsal and ventral setae. Sternite of third 

 pereopods with subquadrate to subrectangular anterior lobe (Fig. 1 b), 

 terminal margin with 3-8 small spines and long setae. 



Male pleopods unknown. Female with pleopods 2-4 well devel- 

 oped, biramous; pleopod 5 appreciably reduced, with rudimentary 

 exopod. Tergite of sixth abdominal somite with deep transverse 

 median furrow; posterolateral margins each with 2 spines, terminal 

 margin with 1 spine one each side of midline. Protopods of uropods 

 (Fig. Id) each with row of small tubercles. Telson (Figs Id, e) with 

 median cleft obsolete or only faintly indicated; terminal margin 

 slightly concave, with row of several large and few smaller spines, 

 extending onto lateral margins. 



COLOUR. Unknown. 



Distribution. At present recognized only from Singapore. 



Etymology. It is with great pleasure that we dedicate this species 

 to Ray Ingle, formerly of the Crustacea Section, the Natural History 

 Museum, in recognition of his many contributions to our knowledge 

 of decapod crustaceans in general, and of the Paguridea of the 

 northeastern Atlantic and Mediterranean regions in particular. 



Affinities. Diogenes inglei sp. nov. most closely resembles D. 

 gardineri in the general shapes of the left and right chelipeds, and in 

 the armature of the pereopods. Diogenes inglei sp. nov. is readily 

 distinguished from Alcock's species by its longer antennal pedun- 

 cles and flagellum which carries a pair of long pinnate setae ventrally 

 on each article. Differences are also apparent in the armature of the 

 chelipeds. However, variation in cheliped armature is common in 

 species of this genus, and our present knowledge of D. inglei sp. nov. 

 is too limited to permit evaluation. 



