40 



p. a. Mclaughlin and p.f. clark 



shorter to nearly equal length of antennular peduncles. Antennal 

 acicle not reaching to distal apex of fourth peduncular segment, with 

 simple or bifid terminal spine, lateral margins usually 1 or 2 spines 

 distally, mesial margins with 3-7 small spines. Antennal flagellum 

 with paired long setae ventrally. 



Left cheliped (Fig. 1 lb) with 1 or more rows of small simple or 

 bifid spines on dorsal margin of merus, ventromesial distal angle 

 with 3 or 4 prominent spines, ventrolateral margin with 2-5 much 

 smaller spines distally; broad upper surface of carpus with 3 rows of 

 small acute or subacute spines, stronger on slightly produced distal 

 angle, outer face angularly convex with tuberculate or spinose 

 protuberance at median distal margin, surface armed with blunt or 

 spinulose tubercles and small spines; lower margin of fixed finger 

 and palm straight, with irregular rows of small tubercles or subacute 

 spines; palm with convex outer surface armed with moderately to 

 closely-spaced tubercles, subacute or acute spines or spinules, and 

 with crest of stronger tubercles or spines proximally near midpoint 

 of proximal margin but not continued to articulation with dactyl, 

 upper margin with irregular usually double row of small spines, 

 strongest on produced upper distal angle; upper surface of dactyl 

 with 3 rows of spines. Right cheliped with noticeable hiatus between 

 dactyl and fixed finger; upper margins of carpus, palm and dactyl 

 each with 1-3 rows of small spines partially obscured by long 

 plumose setae. Ambulatory legs with dorsal margins of carpi each 

 with double row of small spines on dorsal surface of second, usually 

 only single row of smaller spinules on third; propodi with irregular 

 row of small spines or spinules, always on second, frequently on 

 third; mesial faces of dactyls (Fig. 8b) each with 2 rows of rather 

 widely-spaced moderately short setae. Anterior lobe of sternite of 

 third pereopods (Fig. 9b) with roundly rectangular, with tuft of setae 

 on either side of midline. 



Telson (Figs 9c,d) with median cleft; terminal margin of left lobe 

 with 3-6 large spines extending onto lateral margin and several very 

 small spinules medianly, right terminal margin with 4-6 small 

 spines. 



Colour. Carapace rather uniform dark brown. Ocular peduncles 

 uniform light brown, or brown with broad, oblique white band 

 medially; cornea golden. Proximal segment of antennular peduncles 

 dark brown; distal segment with brown chromatophores on white, 

 antennal flagella banded brown and white. Merus and carpus of left 

 cheliped solid light brown, chela white; right cheliped solid light 

 brown. Meri of ambulatory legs white with 2 brown bands; carpi 

 light brown with white band distally; propodi white medially, other- 

 wise dark brown proximally and light brown distally; dactyl dark 

 brown proximally and white distally (after Haig and Ball, 1988). 



Distribution. Indian Ocean, from Red Sea and east coast of 

 Africa to Mergui Archipelago; Malaysia; Vietnam; Philippine Is- 

 lands; Indonesia; New Guinea; northeast coast of Australia. 



Remarks. Lanchester's specimens agree well with Forest's (1956) 

 description and with small specimens of this species from northern 

 Australia, particularly in having spinules on the dorsal surfaces of 

 both the carpi and propodi of the ambulatory legs. This is in contrast 

 to the figures of D. avarus given by Rahayu and Forest (1995: Fig. 

 2 g, h) in which the propodi are unarmed, and the third left pereopod 

 has only a few dorsodistal spines on the carpus. However, we 

 observed a similar lack of propodal armature and reduced carpal 

 spination in specimens from Barunda Beach, Lovina Bali. Rahayu 

 and Forest's illustrated specimen was one of their largest males (SL 

 = 3.5 mm), while our specimens from Bali had SL's of 2.84-2.96 

 mm. The largest of the Lanchester specimens had a SL of only 1 .52 

 mm. It may be that propodal spination is lost with increased animal 



size, as the specimens agree in other morphological characters. The 

 identification of D. avarus by Ajmal Khan and Natarajan ( 1 894: 1 8, 

 fig. 15) is uncertain. 



Diogenes planimanus Henderson, 1893 

 (Figs 4a, 9c,e, 12a) 



Diogenes planimanus Henderson, 1893:416, pi. 39, figs 5, 6; 



Lanchester, 1902:365 (in part); Nobili, 1903a: 15; Dechance, 



1964: 35;Tirmizi and Siddiqui, 1982a:43, figs 21, 22; 1982b:fig. 



18. 

 Diogenes custos var. planimanus: Alcock, 1905b:66, pi. 6, fig. 3; 

 SundaraRaj, 1927:133; Kamalaveni, 1950:80; Gordan, 1956:317. 

 ? Diogenes planimanus: Ajmal Khan and Natarajan, 1984:16. fig. 



13. 



Material examined. Lectotype (herein designated). 9 (SL = 5.7 

 mm) Madras; BMNH 1894:7:21:4. Type locality restricted by 

 lectotype designation to Madras. 



Lanchester material examined. 3 cf, 2 9 (SL = 4.72-6.36 mm) 

 Patani; UMZC, Nov. 30, 1899. 



Diagnosis. Shield (Fig. 4) nearly as broad as long; anterior 





^-^ss^ m^ 



v^t 



1mm 



1mm 



1mm 



1mm 



tlmm T 



Fig. 8 Dactyl of left 3rd pereopod (mesial view), a. Diogenes 



goniochirus Forest. 1956, 9 SL = 2.15 mm, UMZC Nov. 30, 1899; b. 

 Diogenes avarus Heller, 1865,cfSL = 1.53 mm. UMZC Nov. 30, 1899; 

 c. Diogenes planimanus Henderson, 1893, 9 SL = 4.85 mm, UMZC 

 Nov. 30, 1899; d. Diogenes platvoeti nom. nov., holotypecfSL = 4.81 

 mm ZMA De20 1.872; e. Diogenes platvoeti nom. nov., 9 SL = 4.67 

 mm, UMZC Nov. 30, 1899; f. Diogenes mixtus Lanchester, 1902, 

 paralectotypecfSL = 5.81 mm, UMZC 1.10050. 



