46 



p. a. Mclaughlin and p.f. clark 



strong, slender spines; ventrolateral margin with row of acute spines, 

 ventromesial margin with irregular double row of subacute or acute 

 spines, ventral surface spinulose or tuberculate. Ischium with spinules 

 or small spines on ventromesial and ventrolateral margins. 



Right cheliped with moderately long and slender chela; fingers 

 opening nearly horizontally and terminating in strong calcareous 

 claws. Dorsal surface dactyl with 2 rows of spines and 1 additional 

 row on dorsomesial margin, all partially obscured by long setae. Palm 

 with scattered moderately strong spines, partially obscured by long 

 setae, dorsomesial margin with single or row of small spines, mesial 

 face somewhat spinulose; fixed finger with 2 or 3 rows of small 

 spinules and tufts of long setae on dorsal surface; dorsolateral margin 

 not well defined, but with numerous small spinules. Carpus broadly 

 triangular; dorsomesial margin with row of small spines practically 

 obscured by long dense setae, and adj acent row of stronger spines on 

 dorsal surface, dorsolateral margin with single or irregular double 

 row of spines and tufts of setae, distal margin with several spines; 

 lateral face spinulose; mesial face weakly tuberculate. Merus triangu- 

 lar; dorsal margin with row of spinules or small spines and tufts of 

 long setae, 2 or 3 prominent spines at or near distal margin, often 1 

 additional strong spine marginally just laterad of midline; dorso- and 

 laterodistal margins usually with small spinules; lateral face fre- 

 quently with numerous short multifid ridges; ventrolateral margin 

 with row of strong spines or acute spines distally and small, multifid 

 short ridges proximally; ventromesial margin with generally double 

 row of small spinules. Ischium with row of small spines or spinules on 

 ventromesial margin; laterodistal margin with few spinules. 



Ambulatory legs generally similar form left to right; dactyls (Fig. 

 8f) long, approximately 0.20 longer than propodi, slender, curved, 

 slightly twisted; ventral margins each with row of long setae; lateral 

 faces each with longitudinal suture; dorsal margins with small 

 spines becoming obsolete in distal half and long setae; mesial faces 

 each with longitudinal sulcus lined with long setae, row of small 

 spines ventrally decreasing in size and not reaching to distal third 

 and gradually replaced by row of long setae. Propodi each with 2 

 rows of spines on dorsal surface, strongest mesially and separated by 

 flattened, unarmed or intermittently spined longitudinal space; lat- 

 eral face with longitudinal row of spinulose tubercles or spines 

 dorsally and usually numerous simple or multidenticulate tubercles, 

 sometimes only weakly developed; ventral surface faintly spinulose, 

 ventrodistal margin usually with row of denticles extending mesially 

 and laterally; mesial face somewhat spinulose ventrally or with 

 irregular longitudinal rows of small tubercles. Carpi each with row 

 of strong spines on dorsal margin; lateral face with 3-5 usually 

 longitudinal rows of spines, spinules, multidenticulate tubercles or 

 low protuberances; laterodistal margin spinose or spinulose; mesial 

 faces each with longitudinal row of small spines adjacent to dorsal 

 margin (second) or unarmed (third). Meri with dorsal surfaces of 

 second pair distally broadened and armed with irregular double or 

 triple rows of small spines or spinulose tubercles, dorsal margin 

 proximally and on third pereopods each with row of spines; 

 ventromesial margins each with nearly double row of spines or 

 spinulose tubercles; row of small spines on ventrolateral margins; 

 lateral faces, particularly of third weakly spinulose or tuberculate. 

 Sternite of third pereopods subrectangular, with tuft of setae on 

 either side laterally. Sternite of fifth pereopods as slender elongate, 

 tuberculate, calcareous rod. 



Protopod of right uropod with well developed posterior protuber- 

 ance nearly equal to size of endopod, and similarly covered with rasp 

 of corneous scales. Telson (Fig. 9i) without median cleft; terminal 

 margin spinulose on right, strongly spinose approaching left lateral 

 angle and continuing onto lateral margin over approximately half 

 length. 



Colour. Not known. 



Distribution. Malaysia; ? Singapore. 



Remarks. Nobili (1903) reported numerous specimens of D. 

 mixtus collected in Singapore. He distinguished Lanchester's ( 1 902) 

 species from D. intermedius De Man, 1 892 by the presence, in the 

 former species, of spines on the merus of the second and third 

 pereopods. A check of the collections of the Museo Regionale di 

 Scienze Naturali, Torino, failed to locate Nobili's specimens (Elena 

 Gavetti, pers. comm.); therefore it has not been possible to confirm 

 Nobili's (1903) identification. Although it was not represented in 

 their collection, D. intermedius was reported as one of the Indone- 

 sian species of Diogenes by Rahayu and Forest (1995); no mention 

 was made of D. mixtus. 



Paguristes hians Henderson, 1888 

 (Fig. 14) 



Paguristes hians Henderson, 1888:79, p. 8, fig. 4;Alcock, 1905:40, 

 pi. 3, fig. 2; Southwell, 1906:216; Estampador, 1937:506; 

 Thompson, 1943:415; Gordan, 1956:322 (in part) (lit.); Edwards 

 and Emberton, 1980:236 (list); Haig and Ball, 1988: 173; Hogarth, 

 1988:1 100; Morgan, 1990:21. 



Diogenes desipiens Lanchester, 1902:366, pi. 34, figs 1, la;Alcock, 

 1905:165 (list); Gordan, 1956:317 (lit.). 



Non Paguristes hians: Grant and McCulloch, 1906:33; McCulloch, 

 1913:346 = Paguristes monoporus Morgan, 1987. 



Material examined. Holotype. cf (SL = 4.89 mm); BMNH 

 1888.33., H.M.S. 'Challenger' station 208, off Manila. Supplemen- 

 tal material. 1 cf, 2 ovigerous 9 (SL = 1.67-2.51 mm), Maldive 

 Islands, 14 February, 20 December 1993, collector P. Hogarth. 



Diagnosis. Shield (Fig. 1 4) considerably longer than broad. Ros- 

 trum broadly triangular, not produced to level of terminal spinules of 

 obtusely triangular lateral projections. Ocular peduncles long and 

 extremely tenuous, slightly overreaching antennular peduncles and 

 approximately 0.50 longer than antennal peduncles; acicles elongate 

 and nearly rectangular, with large tuberculate terminal spine and 

 smaller accessory spine laterally. Subquadrate calcified lobe with 4 

 subacute spinules on anterior marginal part of, or articulating with, 

 interocular lobes. Antennal acicle long, reaching nearly to mid- 

 length of ultimate peduncular segment. 



Fig. 14 Paguristes hians Henderson, 1888, holotypecf SL = 4.89 mm, 

 BMNH 1888.33, a. whole animal; b. enlarged view of shield and 

 cephalic appendages. 





