ANATOMY AND SYSTEMATICS OF CARDIOLUCINA 



107 



Fig. 15 Geographical distribution of Cardiolucina pisiformis (Thiele). 



DISTRIBUTION. Indonesia and Philippines (Fig. 17). 



Material examined. INDONESIA: Kai Islands, 05°23'S 

 132°37'E, 436-4 13m & other stations in same area between 250- 

 450m (MNHN, Bouchet and von Cosel manuscript). 

 KALIMANTAN: Sibuko Bay, 762 & 589m (USMN 302240). 

 PHILIPPINES: 1 1°58'N 122°02'E, 486-551 m (MNHN) (see Cosel 

 & Bouchet, ms.); off Baliscasag I., S. Bohol, 8 1 2m ( USNM 298742); 

 Iligan Bay, N. Mindanao, 500m (USNM 298616) & 505m (USNM 

 298660); off Claveria, Burias, 390m (USNM 2800546); off N. Ticao 

 I.,420m (USNM 298025); off Ponson I., Camotes Islands, 650m 

 (USNM 298468); off Apo I., S. Negros, 460m (USNM 298759); 

 Sogod Bay, Leyte, 1280m (USNM 298560). 



Cardiolucina rugosa (Hedley, 1909) 



Figs 18-19 



Lucina (Myrtea) seminula Gould, 1 862: 1 74 (non Lucina seminulum 



Deshayes, 1858: 673). 

 Lucina (Codakia) seminula (Gould) -Smith, 1885: 180-181, pi. 13, 



figs 5 & 5a. 

 Phacoides rugosus Hedley, 1909: 427. 

 Lucina seminula Gould - Johnson, 1964: 148, pi. 28 fig. 3. 



Type material. Lectotype selected by Johnson (1964, pi 28 fig. 

 3) (USNM 553). Hedley (1909) introduced the replacement name P. 

 rugosus because L. seminula is preoccupied. The specimen from the 

 type locality illustrated in Fig. 18 was given by Gould to Cuming. 



Type LOCALITY. Hong Kong Harbour, 9-18 m. 



Shell description. Shell solid, small, height to a maximum of 

 3.1mm, subcircular in outline, H/L mean 0.92; moderately inflated, 

 T/L mean 0.38, tumidity to a maximum of 1 .2 mm on a single valve. 

 Shell inequilateral and extended towards anterior. Pronounced pos- 

 terior sulcus with concentric lamellae. Anteriorly, there is a narrow 

 sulcus. Lunule small, heart-shaped. Anterodorsal area shallowly 

 concave with fine concentric lamellae. Escutcheon lanceolate and 

 slightly convex. Exterior sculpture of about 20 thick, slightly 

 recurved, concentric lamellae. Interspaces between concentric la- 



mellae are variable in width. Concentric lamellae often extended 

 into short flanges at margin of posterior sulcus. Radial ribs indis- 

 tinct, flattened and closely spaced. Hinge plate thick. Left valve with 

 single posterior lateral, two cardinal teeth, the anterior of which is 

 thicker and triangular and a single anterior lateral. Right valve with 

 a single posterior lateral tooth, two cardinal teeth, the posterior large 

 and the anterior a thin plate, with a single anterior lateral tooth. 

 Ligament very short, extending from beak to less than half way to 

 posterior lateral tooth. Inner margin with 20-21 sharp crenulations. 

 Small denticles on posterior and anterior dorsal margin. Shallow 

 sinus on posterior margin corresponding to the position of the 

 posterior sulcus. Anterior adductor muscle scar elongate, not but- 

 tressed, detached from the pallial line only at the ventral tip. Posterior 

 adductor muscle scar ovate. Pallial line continuous. Shell colour 

 creamy white. 



Comparison with other species. Cardiolucina rugosa is closely 

 similar to C. semperiana, but it is smaller, less tumid with more 

 prominent, recurved concentric lamellae and greater anterior exten- 

 sion. 



Habitat. 10-250 m with a few dead shells from 540 m, sublittoral 

 silts and muds. 



Distribution. Queensland, Australia to Hong Kong (Fig. 19). 



Material examined. HONG KONG: North Lantau (BMNH; 

 BMNH 1963233; NMW ace. 55. 158; AM C.33806; 9-1 8m (USNM 

 611842, 2 paratypes); (USNM 24252); (USNM 714821). CHINA: 

 off Pratas Islands, 268m (USNM 296955). KALIMANTAN: off 

 Silungan I., 539m (USNM 291027). PHILIPPINES: (USNM 

 21109); Pescador I. Tanon Straits, 412m (USNM 293876); Bohol 

 (BMNH); off Sombrero I.. Balayan, 216m (USNM 312342); S.E. of 

 Bantayan I., 60m (USNM 293115); Destacado I., 218m (USNM 

 298090); off N. Cebu, 125m(USNM 293266); Pujada R., E. 

 Mindanao, 253m (USNM 294765); off Tochanhi Pt, Tawi Tawi 

 Islands, 90m (USNM 292997) & 35m (USNM 283372); 11°43'N, 

 122°34'E, 93-99m (MNHN). AUSTRALIA: Queensland, Cape 

 York (BMNH 1887.2.9.2772-5); Hope Islands, 10-1 8m (BMNH 

 1910.9.28.68-71 and BMNH 1963259); Palm Island (NMW ace. 

 55. 158); Gulf of Carpentaria 16°58'S 140°53'E,9m(AMC015275); 

 Albany Passage, Torres Strait, 9-30m (AM C.36171); Palm Island 

 (AM C.1049); Low Isles, 27m (AM); Cape Sidmouth (AM C. 

 2594); Endeavour Reef, 40m (AM C.44668); Hervey Bay (AM); 

 Barney Point, Port Curtis (AM C.2 1 805); Facing Island (AM C.4428); 

 Quoin Island, Port Curtis ( AM ) ; Northern Territory, Darwin (AM). 

 PAPUA NEW GUINEA: 0.5 mile off Fairfax Harbour, Port Mores- 

 by, 15-18m(AMC309466). 



Cardiolucina semperiana (Issel, 1869) 

 Figs 21-24 



Lucina pisum Reeve 1850 (part), pi. 11, fig. 66b (non L. pisum 



Sowerby 1836). 

 Lucina semperiana Issel, 1869: 82-83. 



Phacoides (Bellucina) semperiana (Issel) - Lamy 1920: 211-213. 

 Dentilucina (Bellucina) macassari Prashad 1932: 163 pi. 5 figs 13- 



16. 

 Lucina semperiana (Issel) - Bouchet & Danrigal 1982: 16, fig. 19. 

 Bellucina semperiana (Issel) - Oliver 1992: 99, pi. 19 fig. 7. 



TYPE MATERIAL. The species concept of C. semperiana was based 

 on the figure in Savigny (1817, pi. 8, fig 12) and one specimen from 

 Suez in the Issel Collection (now lost); the single shell of this species 

 in the Savigny collection (MNHN) illustrated by Bouchet & Danrigal 



