142 



R. bOtTGER-SCHNACK AND R. HUYS 



margin; ventral blade strong and spiniform, with row of setules on 

 posterior surface; dorsal blade strong and broad, with four dentiform 

 processes around distal margin; dorsal elements setiform, the shorter 

 hyaline, flat and densely setose, the longer multiipinnate. 



Maxillule (Fig. 2E) indistinctly bilobed, with numerous spinules 

 on anterior and posterior surfaces. Inner lobe subcylindrical, with 3 

 elements: outermost one spiniform, swollen at base, fringed with 

 coarse spinules, others setiform and bipinnate; innermost one lo- 

 cated along concave inner margin at some distance from other 

 elements. Outer lobe with 4 elements; outermost element spiniform, 

 curved and unipinnate, longer than the following; other elements 

 with sparse spinules, innermost one shortest. 



Maxilla (Fig. 2F) 2-segmented, comprising syncoxa and allobasis. 

 Syncoxa unarmed, surface ornamented with 2 spinular rows and 1 

 large secretory pore. Allobasis produced distally into slightly curved 

 claw bearing 2 rows of very strong spinules along medial margin; 

 outer margin with strong seta almost extending to tip of allobasal 

 claw, ornamented with few strong spinules distally and a thin 

 hyaline lamella bilaterally, tip of seta with tubular extension (Fig. 

 2fl); inner margin with slender pinnate seta and strong basally 

 swollen spine with double row of very strong spinules along the 

 medial margin and single row of shorter spinules along outer margin 

 (Fig. 2f2). 



Maxilliped (Fig. 2G) 4-segmented, comprising syncoxa, basis 

 and 2-segmented endopod. Syncoxa unarmed, without surface orna- 

 mentation. Basis robust, inner margin with 2 spiniform bipinnate 

 elements nearly equal in length; fringe of long pinnules between 

 proximal seta and articulation with endopod; two rows of long 

 setules on anterior surface as illustrated in Fig. 2G. Proximal endopod 

 segment unarmed. Distal endopod segment drawn out into long 

 curved claw, with pinnules along proximal half of concave margin; 

 accessory armature consisting of minute, naked seta on outer proxi- 

 mal margin and unipectinate spine fused basally to inner proximal 

 corner of claw. 



Swimming legs \—4 biramous (Fig. 3A-D), with 3-segmented 

 rami. Spine and setal formulae as follows: 



Coxa 



Basis 



Endopod 



Exopod 



Legl 



0-0 



1-1 



0-1; 0-1; 0,1,5 



1-0; 1-1; 111,1,4 



Leg 2 



0-0 



1-0 



0-1; 0-2; 1,11,3 



1-0; 1-1; 111,1,5 



Leg 3 



0-0 



1-0 



0-1; 0-2; 1,11,2 



1-0; I- 1; 11,1,5 



Leg 4 



0-0 



1-0 



0-1; 0-2; 1,11,1 



1-0; I- 1; 11,1,5 



Intercoxal sclerites well developed, without ornamentation. Coxae 

 and bases of legs 1-3 with surface ornamentation as shown in Fig. 

 3A-C. Bases with plumose (legs 1, 4), short naked (leg 2) or very 

 long naked, outer seta (leg 3); with anterior secretory pore near outer 

 proximal corner; inner portion slightly produced adaxially in legs 2- 

 4 (Fig. 3B-D). Inner basal seta on leg 1 long, spiniform and naked. 

 Respective legs without distinct length differences between rami 

 with endopod slightly longer than exopod. Bases of spines on 

 exopodal and endopodal segments surrounded by small spinules 

 which are most obvious around terminal endopod spines of legs 2 

 and 4 (Fig. 3b,d). Surface ornamentation of all segments with small 

 scales, similar to those on caudal ramus (Fig. IF). 



Exopods. Outer margin of exopod segments with well developed 

 serrated hyaline lamella; inner margin of proximal exopod segments 

 with long setules. Secretory pore present on posterior surface of 

 distal segments. Hyaline lamellae on outer exopodal spines well 

 developed. Terminal spine of leg 1 equal in length to (leg 1) or 

 shorter than (legs 2-4) distal exopod segment. 



Endopods. Outer margin of endopod segments with fringe of long 

 setules. Inner seta of proximal endopod segment slightly swollen. 

 Distal endopod segments with single secretory pore on posterior 

 surface; distal margin of legs 2 and 3 produced into conical projec- 

 tion ornamented with minute spinules anteriorly (Fig. 3B,C,b,c). 

 Outer subdistal spine equal in length to distal spine in legs 2-4. 

 Outer margin of distal segment of leg 1 terminating in a long process 

 obscuring insertion of distalmost inner seta (Fig. 3a). Inner setae of 

 distal endopod segments of legs 2-4 with spinule comb along 

 proximal inner margin; this comb less obvious in leg 1 ; also present 

 on distal inner seta of middle endopod segment in leg 4. 



Leg 5 (Fig. 1G) comprising small plumose seta arising from 

 lateral surface of somite, and small free segment representing exopod. 

 Exopod slightly longer than wide, bearing stout curved seta apically 

 and smaller slender seta inserted subdistally; surface covered with 

 minute denticles 



Leg 6 (Fig. 1C, H) represented by operculum closing off each 

 genital aperture; armed with a spine and a short spinule. 



Egg-sacs paired, oval-shaped; each sac containing approximately 

 40 eggs (diameter 50-65 urn). 



Adult male 



Body length (measured in lateral aspect; from anterior margin of 

 rostral area to posterior margin of caudal rami, calculated as sum of 

 individual somites): 960 pm. Sexual dimorphism in antennule, 

 antenna, maxilliped, legs 5-6, caudal ramus and in genital segmen- 

 tation. 



Proportional lengths (%) of urosomites (excluding caudal rami) 

 9.4 : 70.0.: 3.4 : 3.8 : 3.0 : 10.5 ; proportional lengths (%) of 

 urosomites (caudal rami included) 8.4 : 62.1 : 3.0 : 3.3 : 2.7 : 8.1 : 



1 1.6. Leg 5-bearing somite with transverse row of denticles dorsally 

 ( Fig . 4D) . Caudal rami 1 . 8 times longer than wide , markedly shorther 

 than in female. Caudal setae with proportional lengths as in female. 

 Dorsal surface of genital somite covered with scale-like structures in 

 anterior half, arranged in a specific pattern (not figured). Surface of 

 genital flaps and ventral surface of anal segment ornamented with 

 several rows of small spinules (Fig. 4E). Dorsal surface of caudal 

 ramus covered with minute scales as in female. 



Antennule (Fig. 4B) 4-segmented; distal segment corresponding 

 to fused segments 4-6 of female; relative lengths (%) of segments 

 measured along posterior non-setiferous margin 14.5 : 19.7 : 42.1 : 



23.7. Armature formula: l-[3], 2-[8], 3-[4], 4-[ll+2ae+(l+ae)]. 

 Antenna as in female, except for lateral armature on distal endopod 



segment (Fig. 4G), with third spiniform element being much stouter 

 than in female and shorter than the 2 adjacent setae. 



Maxilliped (Fig. 4C) 3-segmented, comprising syncoxa, basis 

 and 1 -segmented endopod. Syncoxa without surface ornamentation, 

 unarmed. Basis robust, particularly inflated in proximal half form- 

 ing bulbous swelling; anterior surface with 2-3 transverse spinular 

 rows in addition to row of short flat spinules along inner margin 

 (Fig.4C); posterior surface with 3 rows of short spatulated spinules 

 of graduated length along palmar margin (Fig. 4C); with 2 small 

 naked setae inserted within the longitudinal cleft, the proximal one 

 being slightly longer than the distal one. Endopod drawn out into 

 long curved claw, concave margin unornamented; accessory arma- 

 ture consisting of short, unipectinate spine basally fused to inner 

 proximal corner of claw; claw with minute hyaline apex (Fig. 4c). 



Legs 1-4 with armature as in female; variable number of spinules 

 on conical projection of endopods of legs 2 and 3. 



Leg 5 (Fig. 4D, F) exopod not delimited from somite, general 

 shape and armature as in female; small plumose seta arising from 

 lateral surface of somite as in female. 



Leg 6 (Fig. 4E) represented by posterolateral flap closing off 



