40 
M.J. ORLOVA-BIENKOWSKAJA 
cx \\ 
SOK 

Fig. 41S. serrulatus, parthenogenetic female. A — C, E, interpopulational and age variability, A, type series of S. serrulatus var. montenegrinus 
(Montenegro), B, series from the vicinity of Vladivostok, C, type series of S. capensis, E, series from Taimyr, D, head shield, dorsal, F — head, ventral, G, 
head, lateral. 
2nd trunk limb as long as 0.3 and 0.9 or 0.6 and 0.4 of basal segment 
of plumose seta of 1st prominence respectively. Postabdomen with 
9-15 anal teeth on each side. Supra-anal angle rounded. 
Male. Supra-anal angle rounded. Vas deferens opening in middle of 
anal bay. Postabdomen with 3-5 anal teeth on each side. Dorso- 
posterior valve angle with small rounded prominence. There is no 
morphological hiatus between males of S. serrulatus and S. 
semiserratus. The male of S. mirabilis is unknown, so only the 
females of these species are described. 
ETYMOLOGY. The name ‘Coronocephalus’ is derived from the 
words ‘corona’ — ‘crown’ and ‘cephalon’ — ‘head’ and refers to 
spines on the head that are typical of this subgenus. 
REMARKS. The subgenus consists of three species: S. serrulatus, 
S. semiserratus and S. mirabilis sp.nov. The first is distributed 
world-wide. Statistical analysis of its variation (Orlova- 
Bienkowskaja, 1995a) has revealed that it has no geographical 
races and that there is a morphological hiatus between S. 
serrulatus and S. semiserratus in two pairs of independent metric 
characters. In addition, these species differ from each other in the 
number of denticles on the valve margin. S. serrulatus and S. 
semiserratus are sympatric in South America. Therefore, they are 
not subspecies but separate species. S. mirabilis differ from S. 
serrulatus and S. semiserratus in having an elongate ocellus and 
in the absence of denticles on the frons. 
