92 

Fig. 1 Hetereleotris apora, diagram of head in lateral view showing 
positions of superficial neuromasts of lateraosensory system (composite 
based on several specimens from Mauritius). 
SYSTEMATIC ACCOUNT 
Hetereleotris georgegilli sp. nov. 
Figs 2-6 
HOLOTYPE. USNM 344315, 19.7 mm SL female, Mauritius, Flic 
en Flac, 30 m north of entrance to lagoon, 20°16'S 057°22'E, around 
small coral bommie on coral, coral-rock, sand and silt bottom, 4-10 
m, A.C. Gill, D.G. Smith, M.J. Smale, W. Holleman, P. Clark and B. 
Galil, OS May 1995 (field no. PCH 95-M20). 
PARATYPES. Mauritius: BMNH 1997.10.24.1, 1: 20.3 mm SL 
female (subsequently cleared and stained), BMNH 1997.10.24.2, 1: 
22.5 mm SL male, RUSI 56870, 1: 19.8 mm SL female, collected 
with holotype; USNM 344316, 1: 20.7 mm SL male, Albion, off 
Pointe Petite Riviere at end of Avenue Victory, surge area and 
adjacent gutters with sand, pebble and rock bottoms, 0-5 m, A.C. 
Gill, M.J. Smale and W. Holleman, 15 May 1995 (field no. PCH 95- 
M23): USNM 344317, 1: 22.3 mm SL male, Passe de L Ambulante, 
off Le Morne, outside lagoon, 20°26°10"S 057°17’40"E, spur and 
groove with surge, 6-8 m, P.C. Heemstra, A.C. Gill, D.G. Smith, 
TLCP 

Fig. 2 Hetereleotris georgegilli, diagram of head in lateral view showing 
positions of laterosensory pores and superficial neuromasts (composite, 
based primarily on holotype, USNM 344315, and cleared and stained 
paratype, BMNH 1997.10.24.1). Abbreviations: AIOP, anterior 
interorbital pore; AN, anterior nostril; IFP, infraorbital pore; PIO, 
posterior interorbital pore; PN, posterior nostril; PNP, posterior nasal 
pore; POP, postorbital pore; TLCP, terminal lateral canal pore. 
A.C. GILL 
Fig.3  Hetereleotris georgegilli, holotype, USNM 344315, 19.7 mm SL, 
Flic en Flac, Mauritius. 
M.J. Smale, W. Holleman, P. Clark, et al., 18 May 1995 (field no. 
PCH 95-M30). 

DIAGNOSIS. Hetereleotris georgegilliis distinguished from conge- 
ners by the following combination of characters: second dorsal-fin 
rays I,10—11, usually I,10; anal-fin rays I,9; scales ctenoid, restricted 
to posterior part of body and caudal peduncle (behind segmented 
dorsal-fin ray 5—7); and head pores present (posterior nasal, median 
anterior interorbital, posterior interorbital, infraorbital, postorbital 
and terminal lateral canal pores). 
DESCRIPTION. Dorsal-fin rays VI + J,10 (1,10 £4; 1,11 f1); anal-fin 
rays I,9; pectoral-fin pointed with 18/18 (16 f1; 17 £2; 18 f7) rays, the 
lower | (0 f8; 1 £2) ray unbranched, remaining rays branched; upper 
3-5 pectoral-fin rays with free tips; lower pectoral-fin rays slightly 
thickened, more robust than upper rays; pelvic-fin rays I,5; branches 
on first segmented pelvic-fin ray 5/4 (4 f10); branches on second 
pelvic-fin ray 5/5 (4 £4; 5 £5; 6 f1); branches on third pelvic-fin rays 
6/6 (4 f1; 5 f6; 6 f3); branches on fourth segmented pelvic-fin ray 5/ 
5 (3 f3; 4 f4; 5 £3); fifth “segmented’ pelvic-fin ray unbranched, with 
few or no segments, much shorter than other segmented rays 
(subequal to or shorter than spine) and inconspicuous (clearly 
visible only after dissection; Fig. 4); pelvic fins fully separate, 
without connecting membrane or fraenum (Fig. 5); segmented 
caudal-fin rays 9 + 8; branched caudal-fin rays 8+ 8 (7+ 7 f1;8+7 
f4); upper unsegmented caudal-fin rays 5 (4 fl; 5 £4); lower 
unsegmented caudal-fin rays 4 (4 f2; 5 f3); vertebrae 10 + 17; first 
dorsal pterygiophore formula 3-22110; anal pterygiophores preced- 
ing first haemal spine 2; epurals 1. 
Scales ctenoid, restricted to posterior part of body and caudal 
peduncle, extending anteriorly as narrow midlateral wedge or band 
to vertical through second dorsal-fin segmented ray 6/5 (5 5; 6 4; 
7 f1; Fig. 6); lateral scale rows 11/11 (10 f2; 11 £5; 12 f1; 13 £2). 
First gill arch broadly joined to suspensorium by membrane; gill 
opening restricted to pectoral-fin base; branchiostegal rays 5. 
Premaxilla with 3 or 4 irregular rows of conical teeth anteriorly, 



——7 7 Tr 
(pe SSG0RBenccc—a=: 
SRS oO Se 
a 





SR1I+4 
el 
Fig.4 Hetereleotris georgegilli, cleared-and-stained paratype, BMNH 
1997.10.24.1, 20.3 mm SL, ventral view of right pelvic fin and 
basipterygium. Abbreviations: B, basipterygium; SP, spine; SR I—5, 
segmented rays 1-5. Large stipple indicates blue-stained material (see 
text); small stipple indicates interradial membranes. Arrow points 
anteriorly. Scale = 1 mm. 
