208 



G. CSORBA AND P. JENKINS 



9. rostral swelling width (RSW) - the greatest width of the nasal 



swellings; 

 10. median anterior rostral swelling width (MARW) - greatest 

 width in dorsal view. 



To reveal the taxonomic differences and relations between the 

 taxa included in this study the Mann-Whitney U Test and Principal 

 Component Analysis (PCA) were used. For the univariate analysis a 

 non-parametric method was applied since the raw data did not meet 

 the criteria for a normal distribution based on the F-test Both 

 statistical methods were performed by Statistica 5.1, 1984-1995 

 statistical programme of StatSoft Inc. run on a 486 PC. 



Abbreviations used for institutions are: BMNH - The Natural 

 History Museum, London, formerly the British Museum (Natural 

 History); HNHM - Hungarian Natural History Museum, Budapest; 

 MNHN - Museum National d'Histoire Naturelle, Paris; IEBR - 

 Institute of Ecology and Biological Researches, Hanoi. 



RESULTS 



Rhinolophus stheno microglobosus ssp. nov. 



Figs \-A, Table 1 



HOLOTYPE. BMNH 1997.360(fieldnumber9601/Bll),adultmale 

 in alcohol, skull extracted, collected by members of 'Frontier' the 

 Society for Environmental Exploration - Vietnam, between 17 

 January and 18 March 1996. 



Type Locality. Na Hang Nature Reserve, Tuyen Quang Prov- 

 ince, Vietnam, between 22°16' and 22°31'N, 105°22' and 105°29'E, 

 altitude 100-1082 m. Highly diverse evergreen and semi-evergreen 

 primary limestone rainforest (see Hill & Kemp, 1996). 



Paratypes. The same collection data as the holotype: BMNH 

 1997.359 (field number 9601/B10) adult female in alcohol, skull 

 extracted; BMNH 1997.361 (field number 9601/B25) adult male in 

 alcohol, skull extracted; BMNH 1997.362 (field number 960 1/B28) 

 adult female in alcohol; 2 km SE of Pac Ban, Na Hang Nature 

 Reserve, Tuyen Quang Province, Vietnam, 22°19'N, 105°25'E, alti- 

 tude 300 m, 3 March 1997, collected by Gabor Csorba and Pham 

 Due Tien: HNHM 98.1.1. (field number CSOVI 30) adult female, 

 skin, skull and skeleton; HNHM 98.1.2-3. (field number CSOVI 32, 

 33) adult males, in alcohol, skull extracted; IEBR (not catalogued, 

 field number CSOVI 31) adult female, in alcohol, skull extracted. 

 Referred material: Tham Tap Tao, Fang, Chiangmai, Thailand, 

 19°55'N 99°13'E BMNH 1978.974, adult female in alcohol, skull 

 extracted, BMNH 1978.2301, adult skull only; Chanthaburi, Pong 

 Nam Ron, Khao Soi Dao Tai, Thailand, 12°36'N 102°09E 850 m: 

 BMNH 1978.2298-2300, adult skulls only. 



Comparative material. Rhinolophus stheno stheno -West Ma- 

 laysia: Selangor (BMNH 1898.3.13.1 [holotype], 1898.3.13.2-3, 

 1973.606-607); Gunong Benom, Pahang (BMNH 1967.1492, 

 1967.1494, 1967.1497, 1967.1533-1534);BatuPahat,Kangar,Perlis 

 (BMNH 1968.817-818);Tioman Island (HNHM 95.55.2-4); Indo- 

 nesia: Saekaranda, N. E. Sumatra (BMNH 1907.1.9.2, MNHN 

 1903.3); Kalipoetjang, Tji-Tandoei River, Java (BMNH 

 1909.1.5.179-182). 



Rhinolophus malayanus - Thailand: Biserat, Jalor (BMNH 

 1903.2.6.83 [holotype], 1903.2.6.84, 1908.2.5.24-25); Phu Nam 

 Tok, Saraburi (BMNH 1970.1462); Phu Nam Tok Tap Kuang, 

 Khaeng Khoi, Saraburi (BMNH 1978.973); Satun, Muang, Wang 

 Bla Chan (BMNH 1978.2295); Chiangmai, Fang, Tham Tap Tao 



(BMNH 1978. 2296-2297); West Malaysia: Batu Pahat, Kangar, 

 Perlis (BMNH 1968.812); Kisap Forest Reserve, Pulau Langkawi 

 (BMNH 1968.813-816). 



DIAGNOSIS. Anterior median rostral compartments abruptly el- 

 evated but narrow and globular in outline; posterior median rostral 

 compartments very small but slightly inflated dorso-laterally. Skull 

 slender, rostral swelling width < 5.1. zygomatic and mastoid width 

 <9.1. 



Description. A medium-sized horseshoe bat belonging to the 

 ferrumequinum group (sensu Corbet & Hill, 1992), forearm length 

 43.8-47.2, mean 45.46, SD 1.21, n = 8; head and body length 38.8- 

 45.2, mean 43.39, SD 2.73, n = 8; tail length 17.7-23.0, mean 20.0, 

 SD 1 .49, n = 8; hindfoot length 7.8-8.4, mean 8.0, SD 0. 1 8. n = 8; ear 

 length 16.9-18.7, mean 17.89, SD 0.64, n = 8; weight 9-9.5 grams, 

 mean 9.33, SD 0.24, n = 3. Ear medium in length, just reaching the 

 tip of nose when laid forward. Noseleaf with sella almost parallel- 

 sided, only narrowing very slightly, rounded at tip; the connecting 

 process rounded, typical for the ferrumequinum group; the lancet 

 long, straight-sided, its tip cuneate; the supplementary noseleaf 

 clearly visible; the lower lip has three groves (Fig. 1). The dorsal 

 pelage is light yellowish-brown at the base of hairs, reddish cinna- 

 mon-brown above and c. 8 mm long, that of the venter paler and 

 shorter. The wing membranes are uniformly dark brown. The fifth 

 metacarpal is subequal or slightly longer than the fourth, the third 

 shorter than fourth. Ratio of first to second phalange of third digit 

 1.56-1.67, mean 1.62, SD 0.04. n = 8. 



Skull averaging smaller than in R. s. stheno; slender, rostral 

 swelling width less than 5.1 mm, zygomatic and mastoid width 

 subequal, not exceeding 9. 1 mm (see Table 1 ). The anterior median 

 rostral compartments are high and abruptly elevated but narrow and 

 not forming the lateral walls of the rostrum, in profile they are 

 posteriorly concave but less sharply so than in R. s. stheno; the 

 posterior median rostral compartments are slightly inflated dorso- 

 laterally so that the anterior region of the supraorbital depression is 

 shallow and narrow, unlike the deep broad depression of/?, s. stheno; 

 lateral rostral compartments slightly inflated (see Fig. 2). The 

 sagittal crest moderately developed. Palatal bridge less than one- 

 third of the upper toothrow length. Anterior upper premolar well 



Fig. 1 Lateral (left) and frontal (right) views of noseleaves of R. s. 

 microglobosus (HNHM 98.1.2. [paratype]). L = lancet; C = connecting 

 process; S = sella; SN = supplementary noseleaf. Scale = 5 mm. 



