110 



K.L. SANDERS ETAL. 



13 



10 





9? 



A 

 A A A 



5 







AA&^ 



n -j 













A^ 



-5 - 





♦ 





-10 







D n • 



-15 



<$> 





D + 



-20 









-25 



i 



i 



I I 



O Borneo 



A Thailand 



West Malaysia 

 Singapore 



D Siberut 



+ Nias 



% north Sumatra 



<§> central Sumatra 



♦ Betong 



-80 



-60 



-40 



-20 



20 



IU 





■ 



5 



~ 













-5 





^A 



10- 



<§> 





15 



#> 





20 







-20 



O Borneo 



A Thailand 



West Malaysia 

 Singapore 



D Siberut 



+ Nias 



9 north Sumatra 



<§> central Sumatra 



■ south Sumatra 



-15 -10 



10 



15 20 



Fig. 2 Canonical Variate Analysis of T. hageni and T. sumatranus populations (top = males; bottom = females). 





