24 



relative length of the P5 exopod serve to distinguish both species. C. 

 farrani is currently known only from two widely separated localities 

 in the Indo- Pacific, suggesting that it is probably widespread through- 

 out this oceanic basin. 



Clytemnestra longipes sp. nov. 



Type LOCALITY. Great Barrier Reef - see C. farrani sp. nov. 



ETYMOLOGY. The species name is derived from the Latin longus 

 (long) and pes (foot), and refers to the very long male P5 and P6. 



Type material. Holotype 6 in alcohol (BMNH 1999.997). This 

 material was originally registered as C. scutellata under BMNH 

 1948.4.28.121. Collected during Great Barrier Reef Expedition 

 1928-29 on either 15 June (stn 62), 10 July (stn 65) or 18 July 1929 

 (stn 68). 



Description. 



female. Unknown. 



male. Total body length from tip of rostrum to posterior margin of 

 caudal rami: 1211 um. Maximum width (362 urn) measured at 

 posterior margin of cephalic shield. Posterolateral angles of 

 cephalothorax angular, weakly produced (Fig. 14A). Backwardly 

 produced alate processes of somites bearing P2-P4 well developed. 

 Urosome with genital and first abdominal somites separate (Fig. 

 14B). 



Urosomites without dorsal ornamentation; all postgenital somites 

 with multiple rows of minute spinules around ventral rear margin, 

 those on urosomites 3, 5 and 6 arranged in paired patches either side 

 of ventral midline (Fig. 14B). 



Caudal rami (Fig. 14B) with bare seta II and minutely pinnate 

 setae I and III; setae IV-V long (54% of urosome length) and 

 plumose. 



Rostrum (Fig. 14A) rounded anteriorly, protruding. Antennule, 

 antenna, mouthparts (proximal endite on maxillary syncoxa present) 

 and maxillipeds as in type species. 



P2-P4 exp-3 with only 2 outer spines (Fig. 14C). P2-P4 armature 

 formula: 



exopod 



endopod 



P2 

 P3 

 P4 



1.1.222 

 1.1.322 

 1.1.322 



1.2.221 

 1.2.321 

 1.2.221 



P5 (Fig. 14D) narrow and elongate, extending to distal margin 

 of first abdominal somite (Fig. 14B); exopod 2.7 times as long as 

 basis; with 3 outer seta and 1 long seta at apex and subdistal inner 

 corner. 



Sixth pair of legs (Fig. 14E) forming very long cylindrical process 

 with 1 apical and 2 outer bare setae. 



Remarks. The male of this species differs from all known males 

 in (1) the ventral ornamentation pattern of the urosome, displaying 

 spinules on all postgenital somites, and (2) the extreme elongation 

 of the P5 and P6 (the distribution pattern of the 3 elements on the 

 latter indicate that allometric growth must have happened prima- 

 rily in the apical portion of the cylindrical process). C. longipes 

 has the same swimming leg setal formula as C. asetosa but, in 

 addition to the characters listed above, differs from the latter in 

 body size and the presence of the proximal endite on the maxillary 

 syncoxa. 



R. HUYS AND S. CONROY-DALTON 

 Clytemnestra asetosa sp. nov. 



TYPE LOCALITY. Suez Canal. Port Taufiq, Bay of Suez (Egypt). 



ETYMOLOGY. The species name alludes to the absence of the 

 proximal enditic seta on the maxillary syncoxa. 



Type material. Holotype 6 dissected on 10 slides (BMNH 

 1999. 1025). Paratypes in alcohol are 3 99, 2 3 6 ( 1 damaged) and 1 

 cop. V 6 (BMNH 1999.1026-1031); collected during the Cam- 

 bridge Expedition to the Suez Canal, 1924. This material was 

 originally identified as C. scutellata by Gurney (1927) and Boxshall 

 (1979). 



Other material examined. From R. Bottger-Schnack: 3 

 copepodid II stages in alcohol (BMNH 1999.1066-1068); central 

 Red Sea, Meteor cruise 5/5, stn 682 (21°13.9' N, 38°05.7' E); 25 July 

 1987; multiple opening-closing net, 0.055 mm mesh, vertical haul- 

 ing, 10-50 m (total water depth 1890 m). 



Description. 



FEMALE. Total body length from tip of rostrum to posterior margin 

 of caudal rami: 758-830 um (x = 801 pm; n = 3). Maximum width 

 (226 um) measured halfway down the cephalic shield. Posterola- 

 teral angles of cephalothorax rounded, not produced. Backwardly 

 produced alate processes of somites bearing P2-P4 distinctly shorter 

 than in C. scutellata and C. gracilis. General body shape (Fig. 15 A) 

 very similar to that of C. farrani (Fig. 12A). 



Genital double-somite (Fig. 15B) weakly constricted bilaterally; 

 original segmentation marked by minute, paired chitinous patches 

 ventrally. Genital field as in type species. 



Urosomites without dorsal ornamentation; penultimate and anal 

 somites with multiple patches of minute spinules around ventral 

 hind margin (Fig. 15B). 



Caudal rami (Fig. 15C) with bare setae I and II; setae IV slightly 

 shorter than seta V, both plumose. 



Rostrum (Fig. 15 A) rounded anteriorly, not distinctly delimited 

 from cephalic shield. 



Antennule (Fig. 16A) 7-segmented, with reduced armature on 

 segments 2 and 3. Armature formula: 1-[1 plumose], 2-[9 + 1 

 plumose], 3-[3 + 3 plumose + 1 transformed], 4-[l + 1 plumose + (1 

 transformed + ae)], 5-[l], 6-[3], 7-[8 + acrothek]. 



Antenna with weakly defined exopod (Fig. 17G); one seta fused 

 basally to segment. 



Mandible (Fig. 1 6B ). Palp represented by minute seta; gnathobase 

 with large lateral tooth (arrowed in Fig. 16C). 



Maxillule (Fig. 16D) produced into distal lash (derived from 

 armature element); with 1 lateral seta and 1 apical spine. 



Maxilla (Fig. 16E) as in type species except for absence of 

 proximal endite on syncoxa (position in other species arrowed in 

 Fig. 16E). Maxilliped as in C. scutellata. 



PI (Fig. 17 A) as in C. scutellata but setules along inner margin of 

 enp-1 absent. P2-P4 (Fig. 17B-D) with only 2 outer spines on exp- 

 3. P2-P4 armature formula: 





exopod 



endopod 



P2 

 P3 

 P4 



1.1.222 

 1.1.322 

 1.1.322 



1.2.221 

 1.2.321 

 1.2.221 



P5 (Fig. 17E) nearly extending to posterior margin of genital 

 double-somite. Basis short, exopod about 2.5 times as long as basis, 

 with 5 setae (3 outer, 1 apical, 1 inner). 



