58 



R. HUYS AND W. LEE 



seta and 2 small processes; with 4 medially directed, spinous 

 processes (Fig. 2H). 



First postgenital somite with backwardly produced lateral angles, 

 bearing spinular tuft (Fig. 1C); without ventral ornamentation. 

 Penultimate and anal somites distinctly narrower; ventral posterior 

 border with spinules. Anal somite (Figs. ID; 31 A) with spinulose 

 anal operculum. 



Caudal rami (Figs. ID; 31 A) widely separated; slightly longer 

 than widest portion; proximal half distinctly bulbous with major 

 swelling medially, dorsally and ventrally (Fig. 1C); ventral surface 

 with very large semi-circular concavity (Fig. 21) leading to small 

 tube-pore; dorsolateral surface with minute spinules; seta I small, 

 setae II — III well developed, naked and closely set; setae IV and V 

 pinnate and with fracture planes, seta V 2.5 times as long as seta IV; 

 setae VI-VII naked. 



Rostrum (Figs 1A; 2A) large, rounded anteriorly; delimited at 

 base by transverse surface suture; with paired sensillae anteriorly 

 and median tube-pore dorsally. 



Antennule (Fig. 2C) slender, incompletely 7-segmented, with 1 

 minute (obscured by large distal one) and 2 well developed spinous 

 processes on posterior margin of segment 1 , no processes on long 

 segment 2. Segment 1 with short spinules posteriorly between 

 processes and large spinular patch around anterior margin. Armature 

 formula: 1-[1], 2-[4 + 4 pinnate], 3-[6], 4-[(2 + ae)], 5-[l], 6-[2], 7- 

 [6 + 1 pinnate + acrothek]. Aesthetasc on segment 4 fused basally to 

 2 setae. Acrothek consisting of aesthetasc and 2 naked setae; set on 

 apical pedestal. Boundary between segments 6 and 7 only expressed 

 posteriorly. 



Antenna (Fig. 2D) with elongate exopod bearing 2 lateral and 2 

 apical pinnate elements, and a longitudinal row of fine spinules. 

 Allobasis with pinnate abexopodal seta and spinular patch opposite 

 exopod. Endopod with lateral armature consisting of 1 pinnate spine 

 and 2 setae; distal armature consisting of 2 unipinnate spines and 3 

 geniculate setae (outermost fused basally to small tube-seta). 



Labrum (Fig. 2B) with spinules around distal margin; anterior 

 face with dense pattern of fine spinules and distal patch of overlap- 

 ping scales. 



Mandible (Fig. 2E) with short gnathobase and small 1 -segmented 

 palp probably representing fused basis and endopod; with 2 lateral 

 (basal) pinnate setae and 3 distal (endopodal) setae (1 pinnate, 2 

 bare). 



Paragnaths highly ornate lobes as in Fig. 3F 



Maxillule (Fig. 2F) with well developed praecoxal arthrite bear- 

 ing 1 seta on anterior surface and 9 elements around distal margin. 

 Coxal endite with 1 spine and 1 seta, basal endite with 1 spine and 2 

 setae. Exopod a short segment with 2 distal setae; endopod incorpo- 

 rated into basis, represented by 2 setae. 



Maxilla (Fig. 2G). Syncoxa with very long spinules around outer 

 margin and dense surface spinulation as figured; with 3 endites; 

 praecoxal endite small, with 1 plumose seta; middle endite drawn 

 out into pinnate claw, with 2 tube-setae; distal endite with 3 ele- 

 ments. Allobasis produced into strong curved claw; accessory 

 armature consisting of 1 spine and 1 seta; with spinular patch 

 proximal to endopodal setae. Endopod incorporated into allobasis, 

 represented by 2 bare and 2 pinnate setae. 



Maxilliped (Fig.3E) slender, with elongate basis and endopodal 

 claw. Syncoxa with 2 plumose setae. Basis with spinular ornamen- 

 tation as figured; spinules present along entire palmar margin. 

 Endopod represented by very long, minutely pinnate claw bearing 1 

 accessory seta and tube-pore at base. 



PI (Fig. 3 A) with dense ornamentation on praecoxa, coxa and 

 basis. Basis with pinnate seta on anterior surface and along outer 

 margin. Exopod 2-segmented, small compared to endopod; exp-1 



not extending to distal margin of basal pedestal, with pinnate outer 

 spine; exp-2 with 3 pinnate outer setae and 2 geniculate setae 

 apically. Endopod slender; enp-1 about 2.5 times as long as basis, 

 with long setules along inner margin and fine spinules along outer 

 margin; enp-2 about 3 times as long as wide, with slender minutely 

 pinnate claw and small accessory seta (Fig. 3B). 



P2-P4 (Figs 3C; 4A-B) with 3-segmented exopods and 2-seg- 

 mented endopods. P2 basis with long, bipinnate outer spine; P3-P4 

 bases with bare outer seta. P2-P3 enp-1 with multipinnate inner 

 seta; P4-enp-l (Fig. 4C) with basally swollen, minute seta. Outer 

 spine of P2-P4 enp-2 very long and setiform. Tube-pore present 

 near distal outer corner of P3-P4 enp-2. Armature formula as 

 follows: 



Exopod 



Endopod 



P2 

 P3 

 P4 



0.1.123 

 0.1.223 

 0.1.223 



1.221 

 1.321 

 1.221 



[6: 1.1.220] 



P5 (Fig. 3D). Endopodal lobe small, extending just beyond 

 insertion sites of proximal outer setae of exopod; with 1 short and 1 

 long pinnate seta apically, and 2 long widely separated setae along 

 inner margin; tube-pores present near articulation with exopod, 

 between apical setae and proximal to innermost seta. Exopod elon- 

 gate, produced apically into tubular extension bearing 1 bare seta; 

 inner margin with 1, outer margin with 4 pinnate setae; inner seta 

 distinctly longer than apical one. Both baseoendopod and exopod 

 with elaborate ornamentation pattern as figured. 



MALE. Body length from anterior margin of rostrum to posterior 

 margin of caudal rami 5 12 urn (n=2, range 500-524 urn). Maximum 

 width (168 urn) measured at posterior margin of cephalothorax. 



Body (Fig. 5 A) more compact and abbreviated than in 9; covered 

 with similar dense pattern of minute spinules. Pattern of cup-shaped 

 pores as in 9 except for paired lateral pores present on genital 

 somite. Cephalothorax wider than free somites; body constricted at 

 level of genital somite. None of urosomites with backwardly pro- 

 duced posterolateral corners. 



Genital somite with large cup-shaped pores laterally, each partly 

 closed off by fringe of setular extensions (Fig. 5H). Sixth legs 

 represented by well developed opercula, one articulating and clos- 

 ing off left or right genital aperture; each produced into cylindrical 

 process bearing 1 lateral and 1 apical seta. 



Antennule (Fig. 5B-E) 7-segmented, subchirocer, with 

 geniculation between segments 5 and 6. Segment 1 with spinules/ 

 setules around anterior margin and 2 spinous processes along poste- 

 rior margin. Segment 2 longest; segment 4 minute, represented by 

 incomplete sclerite. Segment 5 with large proximal process anteriorly, 

 bearing modified bifid spine (Fig. 5D); forming cylindrical process 

 bearing long aesthetasc. Segment 6 with 3 spinous processes along 

 anterior margin. Distal portion of segment 7 elongated, displacing 

 acrothek to position isolated from other armature. Armature for- 

 mula: 1-[1], 2-[4 + 5 pinnate], 3-[6 + 1 pinnate], 4-[2], 5-[7 + 2 

 pinnate + 1 bifid spine + (2 + ae)], 6-[l + 3 processes], 7-[7 + 

 acrothek]. Apical acrothek consisting of aesthetasc and 2 bare setae. 



P3 endopod (Fig. 4D) 3-segmented; enp-1 as in 9; enp-2 with 

 inner seta and short outer apophysis; enp-3 small, with tube-pore, 2 

 lateral and 2 apical setae. 



P5 (Fig. 5G) medially fused, positioned ventrolaterally. 

 Baseoendopod without endopodal lobe; medial margin with 2 

 spinules and 2 tube-pores; outer basal seta arising from short spinulose 

 pedestal. Exopod free; with 1 apical and 1 inner and 3 outer pinnate 

 setae. 



