82 



R. HUYS AND W. LEE 



P5 9 with separate rami; exopod elongate, with 6 setae/spines; 

 baseoendopod slightly developed, with 5 setae/spines. P5 6 without 

 endopodal lobe; exopod short, with 2 outer, 1 apical and 2 inner 

 elements (distal inner spiniform). Outer basal seta arising from long, 

 articulating, cylindrical setophore in both sexes. 



P6 9forming opercula closing off paired genital apertures; with 2 

 small setae at outer corner. P6 6* asymmetrical; membranous flaps 

 with 1 apical and 1 lateral seta. 



TYPE SPECIES. Laophonte typhlops Sars, 1908 =Archesola typhlops 

 (Sars, 1908) comb. nov. 



OTHER SPECIES. Laophonte longiremis T. Scott, 1905 = A. 

 longiremis (T. Scott, 1905); A. hamondi sp. nov. 



Species INQUIRENDAE. Esola sp. sensu Chislenko (1967); Esola 

 typhlops pontoica Por, 1959 = A. typhlops pontoica (Por, 1959) 

 comb. nov. 



ETYMOLOGY. The Greek prefix arche alludes to the primitive 

 position of the genus. 



Archesola typhlops (Sars, 1908) comb. nov. 



Laophonte typhlops Sars, 1908 



Esola typhlops (Sars, 1908) Lang (1948) 



TYPE LOCALITY. Flekkero, south coast of Norway, 36 m depth. 



Material examined. 



(1) West Runton, Norfolk, England: 1 3 dissected on 8 slides 

 (BMNH 1 999. 1 079); collected among Polyclinum and Morchellium 

 under rocks; leg. R. Hamond, September 1971; 



(2) Frierfjord/Langesundfjord, Norway: 4 damaged 99 (3 in alco- 

 hol: BMNH 1999.1081-1083; 1 dissected on 5 slides: BMNH 

 1999.1080); 99 m, mud, leg. R. Huys, 1985; 



(3) Gullmar Fjord, Sweden: 1 9 in alcohol (NMNH 90955); 30 m, 

 sand; leg. K. Lang, 08 July 1942. 



Redescription. 



FEMALE. Body length from anterior margin of rostrum to posterior 

 margin of caudal rami 585 urn (n=4; range: 575-592 urn). 



Body cylindrical, not dorsoventrally depressed, covered with 

 dense pattern of minute surface ridges dorsally and laterally. 

 Cephalothorax with almost parallel lateral margins in posterior two- 

 thirds, without paired cup-shaped pores. Posterior margin of 

 cephalothorax and all body somites with row of long setules dorsally 

 and laterally. Posterior margin of urosomites with spinules all 

 around (Fig. 23B); ventrolateral areas of cephalic shield and 

 pleurotergites of pedigerous somites with longer setules. Pleurotergite 

 of P5-bearing somite narrowest. 



Genital double-somite (Fig. 23A) wide and dorsoventrally flat- 

 tened; original segmentation marked by bilateral constriction and 

 transverse surface ridge dorsally; without cup-shaped pores in ante- 

 rior half; lateral lobes in both anterior and posterior halves with 

 backwardly directed strong spinules; ventral surface without orna- 

 mentation except for spinules around hind margin and 2 pairs of 

 medial tube-pores. Genital field located near anterior margin (Fig. 

 23A); copulatory pore minute. Sixth legs forming well developed 

 opercula closing off paired genital apertures; each with 2 naked 

 setae. 



Anal somite (Fig. 23B) with coarse spinules on anal operculum. 



Caudal rami (Figs. 23B; 24F; 31C-D) widely separated, cylindri- 

 cal and slightly tapering posteriorly; without cup-shaped pores; 

 about 4 times as long as wide; setae I — III closely set, I minute, II — III 

 very long and thin; setae IV and V pinnate and with fracture planes, 



seta IV distinctly longer than caudal ramus; setae VI-VII naked. 

 Vent-pore and small tube-pore present ventrally near insertion sites 

 of setae I-III (Fig. 31C-D). 



Rostrum as in 6 (Fig. 22B); large, trapezoid with straight anterior 

 margin; delimited at base by transverse surface suture; with paired 

 sensillae anteriorly and median tube-pore ventrally. 



Antennule (Fig. 22A) slender, 7-segmented; segments 1-2 with- 

 out processes. Segment 1 with spinules around anterior margin; 

 segment 4 forming large cylindrical pedestal ventrally. Armature 

 formula: 1-[1], 2-[8 + 1 pinnate], 3-[6], 4-[(2 + ae)], 5-[l], 6- [2], 7- 

 [7 + acrothek]. Aesthetasc on segment 4 fused basally to 2 setae. 

 Acrothek consisting of aesthetasc and 2 naked setae; set on small 

 tubercle. 



Antenna (Fig. 23C) with elongate exopod bearing 2 lateral and 2 

 apical pinnate setae, and a longitudinal row of coarse spinules. Coxa 

 with few large spinules, allobasis with pinnate abexopodal seta. 

 Endopod with lateral armature consisting of 1 seta, 1 large and 1 

 small spine; distal armature consisting of 2 unipinnate spines and 3 

 geniculate setae (outermost fused basally to small seta). 



Labrum as in A. hirsuta. 



Mandible (Fig. 25A) with short gnathobase and small bilobed 

 palp representing partially fused basis and endopod; with 2 lateral 

 (basal) pinnate setae and 3 distal (endopodal) setae; exopod repres- 

 ented by minute segment bearing 1 apical seta. 



Maxillule (Fig. 25B) and maxilla as in E. bulligera. 



Maxilliped (Fig.23D) slender, with elongate basis and endopodal 

 claw. Syncoxa with 2 pinnate setae. Basis with naked palmar margin 

 and setules around outer margin. Endopod represented by very long, 

 naked claw bearing 1 accessory seta at base. 



PI (Fig. 22F) with sparse ornamentation on coxa and basis. Basis 

 with pinnate seta on anterior surface and along outer margin. Exopod 

 3-segmented, well developed; exp-1 with long pinnate outer spine; 

 exp-2 with 1 naked outer spine; exp-3 with 2 unipinnate lateral setae 

 and 2 geniculate setae apically. Endopod long and slender; enp-1 

 with long setules along inner margin and shorter spinules along 

 outer margin, with thin inner seta in distal quarter (arrowed in Fig. 

 22F); enp-2 about twice as long as wide, with strong minutely 

 pinnate claw and small accessory seta. 



P2-P4 as in Sars (1908). P3 enp-2 (Fig. 24B) with setiform outer 

 spine (arrowed). Armature formula typical for genus. 



P5 (Fig. 23E). Endopodal lobe well developed, not extending 

 beyond insertion sites of proximal outer setae of exopod; with 

 distinctly stepped inner margin bearing 2 strong spines and 1 long 

 distal seta (extending beyond apex of exopod); apex with 2 setae, 

 outer one about twice length of inner one; tube-pores present near 

 apical setae and proximal to innermost spine; outer basal seta 

 inserting on cylindrical articulating setophore. Exopod narrow and 

 elongate, produced apically into long tubular extension bearing 1 

 bare seta; inner margin with 1, outer margin with 1 naked and 3 

 pinnate setae. Both baseoendopod and exopod with elaborate or- 

 namentation pattern as figured. 



male. Body length from anterior margin of rostrum to posterior 

 margin of caudal rami 475 urn. Sexual dimorphism in antennule, P3 

 endopod, P5, P6 and genital segmentation. 



Antennule (Fig. 22B-E) 7-segmented, haplocer, with geniculation 

 between segments 5 and 6. Segment 1 with spinules/setules around 

 anterior margin; segment 2 longest; segment 4 minute, represented 

 by incomplete sclerite (Fig. 22C). Segment 5 with large process 

 proximally but forming long cylindrical pedestal distally (Fig. 22D). 

 Segment 6 with 3 spinous processes along anterior margin (Fig. 

 22E). Distal portion of segment 7 elongated, displacing acrothek to 

 position isolated from other armature (Fig. 22E). Armature formula: 



