R. HUYS AND W. LEE 



Rostrum (Fig. 21 A) longer than in A. typhlops; trapezoid with 

 straight anterior and concave lateral margins; delimited at base by 

 transverse surface suture; with paired sensillae anteriorly and me- 

 dian tube-pore ventrally. 



Antennules to maxillipeds as in A. typhlops. 



PI (Fig. 24A) as in A. typhlops except for (a) basal pedestal 

 bearing endopod wider, (b) inner seta on enp-1 inserting more 

 distally, and (c) enp- 1 about twice the length of exopod (distinctly 

 shorter in type species). P2-P4 as in A. typhlops. 



P5 (Fig. 23F). Endopodal lobe well developed, well extending 

 beyond insertion sites of proximal outer setae of exopod; with 

 distinctly stepped inner margin bearing 2 strong spines (more closely 

 set than in A. typhlops) and 1 bare distal seta (not extending beyond 

 apex of exopod); with 2 apical setae, outer one about 1.5 times 

 length of inner one; tube-pores present near apical setae and proxi- 

 mal to innermost spine; outer basal seta inserting on cylindrical 

 articulating setophore. Exopod elongate but distinctly shorter than 

 in A. typhlops, produced apically into short tubular extension bear- 

 ing 1 bare seta; inner margin with 1, outer margin with 4 pinnate 

 setae. Both baseoendopod and exopod with elaborate ornamentation 

 pattern as figured. 



MALE. Unknown. 



REMARKS. Differentiation of A. typhlops and A. hamondi is best 

 achieved by comparison of the general body shape, caudal ramus 

 outline and armature pattern, and 9 P5 morphology and morpho- 

 metry 



Esola sp. sensu Chislenko (1967) 



Chislenko illustrated a male which he obtained in Laminaria 

 saccharina washings from the White Sea and identified as Esola sp. 

 His drawings of the caudal ramus, P3 endopod and PI leave little 

 doubt that this species belongs to Archesola and is obviously close 

 to A. typhlops and A. longiremis. The caudal ramus L:W ratio 

 appears to be intermediate between the latter two species and the PI 

 endopod and exopod have slightly different proportions. Chislenko's 

 male (0.35 mm) is smaller than those of A. typhlops recorded by 

 Drzycimski (1969) from the Bergen area (0.45 mm) and our single 

 male from West Runton (0.475 mm). The antennary exopod bearing 

 the atypical number of 5 setae is obviously based on an aberrant 

 specimen. His illustration of the P3 endopod lacks the proximal 

 inner seta on the distal segment, its location being indicated by the 

 distinct step in the inner margin. Finally, the small inner seta 

 illustrated on the P5 baseoendopod is probably a tube-pore. The 

 White Sea material identified by Brotskaya (1961) as E. longiremis 

 is likely to be conspecific with this species, the true identity of which 

 is as yet uncertain. Consequently, Chislenko's species is tentatively 

 ranked specues inquirenda in Archesola. 



Archesola typhlops pontoica (Por, 1959) comb. nov. 



Esola typhlops pontoica Por, 1959 



Type locality. Black Sea coast, Rumania. 



Type material. Dr Ileana Negoescu (Museum 'Grigore Antipa', 

 Bucharest) informed us that the syntypes no longer exist. 



REMARKS. Por (1959, 1964b) established this subspecies for 3 $2 

 found at 61-69 m depth off the Rumanian coast, however it is 

 doubtful whether his material deserves such status. The author 

 discriminated the Black Sea population on the basis of the presence 

 of 6 setae on the 9P5 exopod (Sars (1908) erroneously figured only 

 5), the slightly shorter caudal rami and the incompletely 3-seg- 

 mented PI exopod (a feature displayed in only 1 specimen!). The 



most significant difference, not mentioned by Por, is found in the 

 proportional lengths of the distal antennulary segments (segments 6 

 and 7 being of equal length). Examination of new material is 

 necessary to resolve the identity of the Rumanian population; E. 

 typhlops pontoica is considered here as subspecies inquirenda. 



Genus Corbulaseta gen. nov. 



The diagnosis below is based on Vervoort's (1964) redescription of 

 E. bulligera and personal observations of Wells' (1970) material 

 from Great Britain Rock, Isles of Scilly, and additional specimens 

 collected from the Belgian North Sea coast by the senior author. 



Diagnosis. Laophontidae. Body cylindrical; posterolateral cor- 

 ners of 9 genital double-somite and second abdominal somite 

 laterally and backwardly produced. Integument of cephalothorax 

 and body somites with dense pattern of spinules and setules. Ros- 

 trum large, partly delimited at base by incomplete surface furrow. 

 Cephalothorax with one pair of large, anterodorsal cup-shaped 

 pores; such pores absent on genital (double-)somite and caudal rami. 

 Anal operculum spinulose. Caudal rami rectangular, short; not 

 sexually dimorphic. 



Sexual dimorphism in antennule, P3 endopod, P5, P6 and in 

 genital segmentation. 



Antennules slender; 6-segmented in 9, subchirocer and 7-seg- 

 mented in 6; segment 1 with 1-2 minute processes along posterior 

 margin; with aesthetasc on segment 4 (9) or 5 (6) and as part of 

 apical acrothek on distal segment; segment 5 3 swollen, without 

 anterior outgrowth; proximal aesthetasc fused basally to 2 setae. 

 Antenna with 4 setae on exopod; allobasis with abexopodal seta. 

 Labrum with distal spinular ornamentation. Mandible with 1 -seg- 

 mented palp; exopod and endopod represented by small tubercles 

 bearing 1 and 3 setae, respectively; basis represented by 2 apical 

 setae. Maxillule with minute, defined exopod. Maxilla with 3 endites 

 on syncoxa; endopod represented by 3 setae. Maxilliped slender; 

 syncoxa with 2 setae; entire palmar margin with long setules; 

 endopodal claw elongate. 



PI with 2-segmented exopod bearing 5 setae on exp-2 and 

 elongate endopod; enp-1 without inner seta, enp-2 with minute seta 

 and long, slender claw. P2-P4 with 3-segmented exopods and 2- 

 segmented endopods. P2 basis with short outer spine. Outer spine of 

 P2-P4 enp-2 setiform and very long in P3-P4. P4 endopod modified 

 in both sexes; distal inner seta proximally dilated, bearing enlarged 

 spinules which enclose long secretory tube-pore arising from seg- 

 ment. P3 endopod 3 3-segmented; enp-2 with inner seta and short 

 outer spinous apophysis. Armature formula as follows: 



Exopod 



Endopod 



P2 



P3 



P4 



0.1.123 



0.1.223 

 0.1.223 



1.221* 

 1.321 



0.221 



[6: 1.1.220] 



*: or 1.220 in Vervoort's (1962) 9 specimen of E. bulligera from New Caledonia. 



P5 9 with separate rami; exopod elongate, with 6 setae/spines; 

 baseoendopod slightly developed, with 4 setae/spines. P5 6 without 

 endopodal lobe; exopod short, with 1 inner, 2 apical and 2 outer 

 setae/spines. 



P6 9 forming opercula closing off paired genital apertures; with 2 

 small setae at outer corner. P6 6* asymmetrical; membranous flaps 

 with 1 long and 1 minute seta. 



