102 



ESOLINAE subfam. nov. 



R. HUYS AND W. LEE 



LAOPHONTINAE 



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Fig. 32 Strict component consensus tree of 84 MPTs produced by parsimony analysis. Numbers refer to apomorphic character states listed in Table 3 (15 1 

 and 15 2 denote multistep states). 



antennulary segment 4 (or 5 in cT) provide overwhelming support for 

 the monophyletic status of its sistergroup comprising Mourephonte 

 and 'Esola ' sensu lato. 



The phylogenetic analysis unequivocally identifies the paraphyly 

 of the genus Esola (as originally and pre-cladistically conceived). 

 Three northwestern European species and the unidentifiable Esola 

 spec, sensu Chislenko (1967) form a basal monophyletic group 

 (Archesola gen. nov.) defined by the 2-segmented 3 P3 endopod and 



the presence of an articulating basal setophore on the fifth legs of 

 both sexes. The degree of resolution within this clade will undoubt- 

 edly increase upon the discovery of the males of A. hamondi and A. 

 longiremis. 



Evolution in the outgroup of Archesola is marked by a stepwise 

 addition of modified integumental pores. Initially, only paired 

 anterodorsal (or -lateral) pores were present on the cephalothorax (in 

 compacta, bulligera and possibly spelaea). This condition was 



