260 Variation in Aistivation. [May, 
Calling the outer petal number one, the next outside number 
two, and so on, I can show their position as follows by numbers 
which are here shown as we come to them passing around the 
flower to the right : — 
In 2 flowers the petals formed a spiral to the right.............-. 1, 2, 3, 4, 5. 
irae“ i A ota dette meki ga 1, 5, 4, 3, 2. 
In 5 & ** stood thus 1, 4,.5,; 3,2. 
Bt ee Soa “ were quincuncial to right 1, 4, 2, 5, 3. 
In 3 "s TRN e T rere 1, 3, 5, 2, 4. 
In 3 af at EUG £U0 FER « dave eel E 1, 2, 4, 5, 3. 
In 3 zy = R bts TE E Cheeta ds Been 1, 3, 5, 4, 2. 
: Re EE DROOL PEERS 1, 5, 4, 2, 3. 
In 2 iy A E A AA E a 1, 3, 2, 5, 4. 
In 1 flower n TEENS aT AT 1, 2, 5, 4, 3. 
In 1 4 Tears TE EEN E viv E N pede 1, 4, 3, 5, 2. 
Se ee a, Se a 1, 2, 3, 5, 4 
In 1 . miea e E 1, 3, 4, 5, 2. 
nro “ a EAE JE Severe 1, 5, 4, 3, 2. 
mr E pi petals. wenua aka oao RRN 1, 4, 3, 2. 
In 1 s “ “ “ce 1, 3, 4, 2 
In Agrimonia it was very easy to determine the relative posi- 
tion of each sepal and petal, and also the same with reference to 
the axis. Taking all these things into consideration, of sixteen 
buds examined closely only two were alike. Undoubtedly there 
were still many other variations. The five sepals in each case 
were quincuncial ; in eight the spiral went to the right ; in eight 
the spiral went to the left. In all cases observed the second 
sepal was next to the axis. Of the petals two were arranged in 
quincunx: with one of these the spiral turned to the right; with 
the other to the left. The petals of one: were convolute; one 
spirally imbricated to the right, thus, 1, 2, 8, 4, 5. The rest 
were imbricated in various other ways. 
Dr. Gray, Bentham and Hooker, and others speak of the petals 
of the Onagracee as convolute (contorta) in the bud. The ma- 
jority of cultivated fuchsias are not conyolute in æstivation.? 
Of five hundred and seventy-nine buds of Epilobium angustifo- 
lium the petals of two hundred and eighty-five are conyolute, 
turning to the left, and one hundred and fifty-four to the right. 
Fifty-three had a single spiral turning to the left, and fifty-three 
to the right. In twenty-three, two opposite petals covered both 
edges of the other two. In eight flowers one petal was exterior, 
one interior, and the other two petals of course were alike half- 
covered: and half-covering. In three flowers one petal was out- 
side and the other three were convolute. ; 
1 See note in American Naturalist, page 110, 1876. 
