Bull. not. Hist. Mus. (Zool.) 59(1): 45-81 Issued 24 June 1993 



A revision of Danielssenia Boeck and Psammis 

 Sars with the establishment of two new genera 

 Archisenia and Bathy psammis (Harpacticoida: 

 Paranannopidae) 



RONY HUYS* 



Department of Zoology, The Natural History Museum, Cromwell Road, London SW7 5 BD 



J. MICHAEL GEE 



Plymouth Marine Laboratory, Prospect Place, The Hoe, Plymouth PL1 3DH 



CONTENTS 



Introduction 46 



Materials and Methods 46 



Systematics 47 



Family Paranannopidae Por, 1984 47 



Genus Archisenia gen . nov 47 



Archisenia sibirica (Sars, 1898) comb, nov 47 



Remarks 51 



(i) Synonymy 51 



(ii) Autapomorphies 60 



(iii) Intersexuality 60 



Genus Danielssenia Boeck , 1872 62 



(i) Danielssenia fusiformis (Brady , 1880) nee Sars (1910) 62 



(ii) Danielssenia robusta Sars, 1921 62 



(iii) Danielssenia similis Chislenko, 197 1 64 



(iv) Danielssenia typica Boeck, 1872 64 



( v) Amended diagnosis 67 



Genus Bathypsammis gen. nov 67 



Bathypsammis longifurca (Bodin, 1968) comb, nov 69 



Genus Psammis Sars, 1910 72 



(i) Psammis borealis Klie , 1 939 72 



(ii) Psammis kliei : Smirnov, 1946 72 



(iii) Psammis longipes Becker, 1974 72 



(iv) Psammis longisetosa Sars, 1910 72 



(v) Amended diagnosis 75 



Discussion 78 



Key to genera of Paranannopidae 79 



Acknowledgements 80 



References 80 



Synopsis. Archisenia gen. nov. is proposed to accommodate the sibirica-gvoup of the genus Danielssenia Boeck, 

 1872. Re-examination of Alaskan material of D. stefanssoni Willey, 1920 has shown the latter species to be a junior 

 synonym of D. sibirica Sars, 1898, the type and only species of the new genus. 



Danielssenia robusta Sars, 1921 and Fladenia intermedia (Wells, 1965) are synonymous and consequently F. 

 robusta comb. nov. becomes the type species of the genus Fladenia Gee & Huys, 1990. Danielssenia similis 

 Chislenko, 1971 is regarded as species inquirenda and the genus Danielssenia is redefined from the type species D. 

 typica Boeck, 1872, and two other species (D. quadriseta Gee, 1988 and D. reducta Gee, 1988). 



The status of D. fusiformis (Brady, 1880) nee Sars (1910) is reconsidered and as a result the genus Sentirenia Huys 

 & Gee, 1992 is relegated to a junior synonym of Jonesiella Brady, 1880 which is reinstated to accommodate J. 

 fusiformis Brady, 1880 and J. eastwardae (Coull, 1971) comb. nov. 



Psammis borealis Klie, 1939 is removed from the genus Psammis Sars, 1910 but retained in the Paranannopidae as 

 species incertae sedis. P. longifurca Bodin, 1968 is transferred from Psammis to Bathypsammis gen. nov. The genus 

 Psammis is redefined on the basis of the type species P. longisetosa Sars, 1910, and P. longipes Becker, 1974. 



* Visiting Research Fellow of the Institute of Zoology, University of Gent, B-9000 Gent, Belgium. 



