64 



unknown at that time he regarded such an allocation as being 

 premature. We have re-examined Sars' material of this 

 species from RisOr, Norway (13 9 $ and 1 copepodid V stage; 

 Zoologisk Museum, Oslo, Reg. No. F20257) and found the 

 following significant discrepancies from the original descrip- 

 tion of Sars (1921): (i) there is an inner seta on exp-1 of 

 P2-P4; (ii) the inner element on enp-1 of P2-P4 is a pinnate 

 spine; (iii) the P5 baseoendopod has 4 setae, the second inner 

 one being much smaller than the others; (iv) the P5 exopod is 

 fused to the baseoendopod on the posterior surface. Further, 

 we have made a detailed comparison of these females with 

 the recently discovered female of D. intermedia (which was 

 assigned to the genus Fladenia by Gee & Huys (1990)) and 

 have found them to be identical. Therefore D. robusta must 

 be referred to the genus Fladenia whose type species now 

 becomes F. robusta (Sars, 1921) comb. nov. as this has 

 priority over F. intermedia (Wells, 1965). F. robusta has also 

 been recorded from the Mediterranean by Por (1964), who 

 found one female in 470 m off the coast of Israel, and Soyer 



(1970) who found 18 adult females at depths ranging from 50 

 to 420 m in the vicinity of Banyuls-sur-mer. Both authors 

 state in their text that these specimens agree exactly with the 

 original description. However, Por's (1964) Fig. 73 does show 

 an inner seta on exp-1 of what is probably P4 (this limb is 

 labelled PI by Por but cannot possibly be so as the endopod is 

 3-segmented). This, taken in conjuction with his figure of the 

 P5 (1964, Fig. 74) leaves little doubt that the Mediterranean 

 material can be assigned to F. robusta, thus giving this species 

 a Boreo-Mediterranean distribution similar to that of Jone- 

 siella fusiformis (see Huys & Gee, 1992). 



(iii) Danielssenia similis Chislenko, 1971 



Chislenko (1971) distinguished D. similis from D. typica on 

 the basis of the following characters: (i) Size, the specimen 

 drawn in his Fig. 1 is approximately 0.9 mm long; (ii) a 

 maxilliped with only 1 seta on the syncoxa and a somewhat 

 longer seta on the basis; (iii) the sexual dimorphism on P2 

 endopod, with the loss of the inner seta on enp-1 and of 1 seta 

 on enp-3. The character of size is of no particular significance 

 as it is within (but near the upper limit of) the size range of D. 

 typica given by Gee (1988b). Similarly, the absence of a large 

 seta on the basis of the maxilliped is of doubtful significance 

 as this seta can be easily dislodged during dissection, as was 

 the case in Sars' (1910) description of D. typica (see Gee, 

 1988b). The differences in sexual dimorphism of the male P2 

 endopod are more difficult to assess from drawings alone. 

 However, it is highly improbable that the inner seta on P2 

 enp-1 is missing in the male when it is present in the female as 

 this condition is found in no other member of this genus or 

 indeed of the family as a whole. The same goes for a 

 reduction in the number of setae on enp-3. In Danielssenia, 

 the 2 terminal setae on this segment are very reduced and 

 implanted close together and it is conceivable that Chislenko 



(1971) has combined these 2 fine setae and drawn them as 

 one broad one. We believe that D. similis is referable to D. 

 typica which has been shown to be the most variable species 

 in the genus (Gee, 1988b) but without being able to examine 

 topotype material we must regard it as a species inquirenda. 



(iv) Danielssenia typica Boeck, 1872 



The following material of the Norman collection (The Natu- 

 ral History Museum) has been examined (species name given 



R. HUYS AND J. M. GEE 

 on the original museum label presented in parentheses): 



1911.11.8.43451-470: vial containing > 400 specimens, 

 mostly 99> a gift from T. Scott; collected near Duke Buoy, 

 Plymouth, 01 August 1889; 



1911.11.8.43471-490: vial containing 23 99 and 1 Q\ a gift 

 from T. Scott; collected from Varanger Fjord, East Finmark, 

 Norway, 1890; 



1911.11.8.43491-510: vial containing 31 99 and 3 cfd\ a 

 gift from T. Scott; collected from Vadso, East Finmark, 

 Norway, 03 July 1890; 



1911.11.8.43511-530: vial containing 39 specimens (D. 

 typica), a gift from T. Scott; collected in Trondhjem Fjord, 

 Norway, 1893; 32 9$ belong to D. typica, the other 7 $$ 

 belong to two different species of Halectinosoma; 



1911.11.8.43531-540: vial containing 16 specimens (D. 

 typica), a gift from T. Scott; collected from Inchkeith in Firth 

 of Forth, October 1895. None of these specimens belongs to 

 D. typica, instead the vial contained Bradya sp. (2 99' 8 

 copepodids), 2 cfcf Robertsonia tenuis (Brady & Robert- 

 son), 1 9 Idomene coronata (T. Scott) and 3 99 OI a 

 Fladenia-Uke paranannopid; 



1911.11.8.43541-560: vial containing > 1000 specimens, 

 mostly 99- a gift from T. Scott; collected from Karnes Bay, 

 Isle of Cumbrae, 1888; a second lot of about 200 specimens 

 from the same locality is registrated under no. 1900.3.29.274; 



191 1.11. 8. M. 2299: 1 9 dissected on slide (Jonesiella spinu- 

 losa), dried out; collected in Trondhjem Fjord, Norway, 

 1893; 



1911. 11. 8. M. 2301: 43 specimens mounted in toto on slide 

 (Jonesiella spinulosa), dried out; collected near Duke Buoy, 

 Plymouth, 02 August 1889; 



1911. 11. 8. M. 2300: 8 specimens mounted in toto on slide 

 {Jonesiella spinulosa), dried out; collected from Vadso, East 

 Finmark, Norway, 1890; 



1900.3.6.644: 5 99 mounted in toto and 3 99 (one 

 belonging to Halectinosoma sp.) dissected on slide (Jonesiella 

 spinulosa); collected in Trondhjem Fjord, Norway, 1893. 



Gee's (1988b) redescription of D. typica is updated here by 

 the following observations and illustrations (Figs 14-16) 

 based on specimens from Duke Buoy (closest to type local- 

 ity): 



Somatic hyaline frills of pedigerous and abdominal somites 

 minutely dentate (Fig. 14A) except for the dorsal frill of 

 P5-bearing somite which is deeply incised, forming rectangu- 

 lar lappets (Fig. 14A, B). Frill of cephalothorax smooth. 

 Dorsal transverse spinule rows are found only on thoracic 

 somites bearing P3-P5, the genital double-somite and second 

 abdominal somite. Genital double-somite with continuous 

 transverse chitinous rim dorsally, laterally and ventrally, 

 marking original segmentation (Figs. 14A, D; 16D-E). 

 Pseudoperculum (Figs. 14E-F) formed by deeply incised 

 posterior extension of penultimate somite. Pattern of caudal 

 rami setae as in Figs. 14E-F. 



Rostrum (Fig. 161) large, hyaline, with 2 pairs of minute 

 sensillae; typically deflected (Figs. 14A-C). 



Male antennule (Fig. 15G) 8-segmented or indistinctly 

 9-segmented; distal 2 segments very small and largely fused. 



Mandible with blunt teeth and a single pinnate seta on 

 gnathobase (Fig. 15 A). Palp with short, equally long, 

 1-segmented rami (Fig. 15B); basis with row of very long 

 setules proximally, inner margin with 1 short and 2 long 

 setae; endopod with 2 lateral and 6 apical setae; exopod with 



