80 



R. HUYS AND J.M. GEE 



P2 enp-2 with 1 inner seta 



10. 



REFERENCES 



9. Caudal rami 5 times as long as maximum width; PI endopod 

 shorter than exopod; P5 $ with fused exopod and baseoen- 



dopod Bathypsammis gen. nov. 



Caudal rami broader than long; PI endopod longer than exo- 

 pod ; P5 $ with separated exopod and baseoendopod 



Jonesiella Brady, 1880. 



10. Body dorsoventrally flattened; caudal rami setae IV and V 

 stubby and spiniform; PI enp-1 1.5 times as long asenp- 



2 Peltisenia Huys & Gee, 1993. 



Body not dorsoventrally flattened; caudal rami setae IV and V 

 long and setiform; PI enp-1 at most as long as enp-2 11. 



11. Antennule $ 4-segmented; club-shaped aesthetascs present on 

 mandible (endopod), maxillule (basis) and maxilla (endopod); 

 P2 enp-2 d" without distinct outer apophysis 



Paradanielssenia Soyer, 1970. 



Antennule 9 6-segmented; no club-shaped aesthetascs on 

 mouthparths; P2 enp-2 d* with long outer apophysis 12. 



12. Antennary exopod with 1 seta on proximal segment; P3 exp-3 

 with 7 setae/spines; P2 enp-3 with inner distal seta transformed 

 into large pinnate spine reaching beyond apophysis of enp- 



2 Afrosenia Huys & Gee, 1993. 



Antennary exopod with 2 setae on proximal segment; P3 exp-3 

 with 8 setae/spines; P2 enp-3 d with inner distal seta not 

 transformed and shorter than apophysis of enp-2 



Archisenia gen . nov . 



13. P2 enp-2 with 2 inner setae Psammis Sars, 1910. 



P2 enp-1 with 0-1 inner setae 14. 



14. Club-shaped aesthetascs present on mandible (endopod), max- 

 illule (basis) and maxilla (endopod); P2 exp-3 with at most 6 



setae/spines 15 . 



No club-shaped aesthetascs present on these appendages; P2 

 exp-3 with 7 setae/spines Fladenia Gee & Huys, 1990. 



15. PI enp-2 with 2 terminal setae geniculate; P5 $ baseoendopod 

 and exopod indistinguishable, with 5 setae; P2 enp-2 d" without 



apophysis; P6 c? with 2 setae 16. 



PI enp-2 with 1 terminal seta geniculate; P5 9 baseoendopodal 

 and exopodal lobes indistinguishable, with 3 and 4 setae, 

 respectively; P2 enp-2 cf with small apophysis; P6 d" with 3 

 setae Micropsammis Mielke, 1975. 



16. Antennule in both sexes with densely opaque, bulbous append- 

 age on distal segment P2-P4 exp-2 without inner seta 



Leptotachidia Becker, 1974. 



Antennule in both sexes without densely opaque, bulbous 



appendage on distal segment P2-P4 exp-2 with inner seta 



Telopsammis Gee & Huys, 1991. 



Acknowledgements. The authors wish to thank Dr. Philippe Bodin 

 and the Curators of Crustacea at the Zoologisches Museum der 

 Universitat Kiel, the Naturhistoriska Riksmuseet Stockholm, the 

 Zoologisk Museum Oslo and the National Museum of Natural 

 History (Smithsonian Institution) for the loan of material. Dr Geof- 

 frey A. Boxshall is acknowledged for his comments on an earlier 

 draft of the ms. For the senior author this is communication No. 551 

 of the Centre for Estuarine and Coastal Ecology, Yerseke and for the 

 junior author the work forms part of the Community Ecology 

 Programme of the Plymouth Marine Laboratory. 



Arlt, G. 1983. Taxonomy and ecology of some harpacticoids (Crustacea, 



Copepoda) in the Baltic Sea and Kattegat. Zoologische Jahrbiicher Systema- 



tik. Abteilung fur Systematik 110: 45-85. 

 Becker, K.-H. 1974. Eidonomie und Taxonomie abyssaler Harpacticoida 



(Crustacea, Copepoda). Teil I. Cerviniidae - Ameiridae. 'Meteor- 



Forschungs-Ergebnisse Reihe D 18: 1-28. 

 Bodin, P. 1968. Copepodes Harpacticoides des etages bathyal et abyssal du 



golfe de Gascogne. Memoires. Museum national d'Histoire naturelle. Nou- 



velle Serie (Zoologie) 55: 1-107. 

 Brady, G.S. 1880. A monograph of the free and semi-parasitic Copepoda of the 



British Islands. Vol. 2. 182 pp. Ray Society, London. 

 & Robertson, D. 1876. Report on dredging off the coast of Durham and 



North Yorkshire in 1874. Report of the forty-fifth meeting of the British 



Association for the Advancement of Science 45: 185-199. 

 Bremer, H. 1914. Zwei Falle von Pseudohermaphroditismus bei Diaptomus 



vulgaris Schmeil. Zoologischer Anzeiger 44: 572-574. 

 Cattley, J.G. 1949. Effects of parasitism on the sex of copepods. Report. 



Challenger Society for the Promotion of the Study of Oceanography 3: 11-12. 

 Chislenko, L.L. 1971. Novye massovye formy garpaktitsid (Copepoda, Harpac- 

 ticoida) iz zaliva Pos'eta Yaponskogo morya [New common forms of 



harpacticids (Copepoda, Harpacticoida) from Possjet Bay of the Sea of 



Japan], lssledovaniya Fauny Morel 



8(14): 151-181. [In Russian.] 

 Coker, R.E. 1938. A copepod Intersex'. Travaux de la Station zoologique de 



Wimereux 13: 97-100. 

 Conover, R.J. 1965. An intersex in Calanus hyperboreus. Cruslaceana 



8:153-158. 

 Francois, Y. 1949. Quelques cas d'intersexualite chez Eudiaptomus gracilis 



(Sars) (Copepode Calanoide). Etude morphologique. Bulletin de la Societe 



zoologique de France 74: 232-239. 

 Gee, J.M. 1988a. Some harpacticoid copepods (Crustacea) of the family 



Tachidiidae from sunlittoral soft sediments in Norway, the Celtic Sea and 



Gulf of Mexico. Zoologica Scripta 17: 181-194. 

 1988b. Taxonomie studies on Danielssenia (Crustacea. Copepoda, Har- 

 pacticoida) with descriptions of two new species from Norway and Alaska. 



Zoologica Scripta 17: 39-53. 

 & Huys, R. 1990. The rediscovery of Danielssenia intermedia Wells. 1965 



(Copepoda, Harpacticoida): a missing link between the 'danielsseniid' 



genera and Paranannopus Lang. 1936 (Paranannopidae). Journal of Natural 



History 24: 1549-1571. 

 & 1991. A review of Paranannopidae (Copepoda: Harpacticoida) 



with claviform aesthetascs on oral appendages. Journal of Natural History 



25: 1135-1169. 

 Huys, R. & Boxshall, G.A. 1991. Copepod Evolution. 468 pp. Ray Society, 



London. No. 159. 

 & Gee, J.M. 1990. A revision of Thompsonulidae Lang, 1944 (Copepoda: 



Harpacticoida). Zoological Journal of the Linnean Society 99: 1-49. 

 & 1992. A revision of Danielssenia perezi Monard, D. paraperezi 



Soyer, D. eastwardae Coull (Harpacticoida: Paranannopidae) and their 



transfer to a new genus. Zoological Journal of the Linnean Society 104: 



31-56. 



& In press. Three new genera of Paranannopidae (Copepoda: 



Harpacticoida). Journal of Natural History: in press. 



& Thistle, D. 1989. Bathycamptus eckmani gen. et spec. nov. (Copepoda, 



Harpacticoida) with a review of the taxonomie status of certain other 

 deepwater harpacticoids. Hydrobiologia 185: 101-126. 



Ianora, A., Mazzocchi, M.G. & Scotto di Carlo, B. 1987. Impact of parasitism 

 and intersexuality on Mediterranean populations of Paracalanus parvus 

 Copepoda, Calanoida). Diseases of Aquatic Organisms 3: 29-36. 



Jespersen, P. 1939. Investigations on the copepod fauna in East Greenland 

 waters. Meddelelser om Gr0nland 119 (9): 1-106. 



Klie, W. 1939. Diagnosen neuer Harpacticoiden aus den Gewassern um Island. 

 Zoologischer Anzeiger 126: 223-226. 



1941. Marine Harpacticoiden von Island. Kieler Meeresforschungen 5: 



1-44. 



1944. Ein gynandromorpher Amphiascus (Cop. Harp.) von Helgoland. 



Zoologischer Anzeiger 145: 77-79. 



Lang, K. 1936a. Undersokningar over Oresund. Untersuchungen aus dem 



Oresund. XX. Harpacticiden aus dem Oresund. Acta Universitatis Lunden- 



sis., n. ser., Avd. 2, 31: 1-52. 

 1936b. Die wahrend der schwedischen Expedition nach Spitzbergen 1898 



und nach Gronland 1899 eingesammelten Harpacticiden. Kungliga Svenska 



Vetenskapsakademiens Handlingar (3)15(4): 1-55. 



1944. Monographic der Harpacticiden (Vorldufige Mitteilung). 39 pp. 



Almqvist & Wiksell, Uppsala. 



