The Bulletin of The Natural History Museum (formerly: Bulletin of the British Museum 

 (Natural History)), instituted in 1949, is issued in four scientific series, Botany, 

 Entomology, Geology (incorporating Mineralogy) and Zoology. 



The Zoology Series is edited in the Museum's Department of Zoology 

 Keeper of Zoology: Dr C.R. Curds 



Editor of Bulletin: Dr N.R. Merrett 



Assistant Editor: Dr B.T. Clarke 



Papers in the Bulletin are primarily the results of research carried out on the unique and ever- 

 growing collections of the Museum, both by the scientific staff and by specialists from elsewhere 

 who make use of the Museum's resources. Many of the papers are works of reference that will 

 remain indispensable for years to come. All papers submitted for publication are subjected to 

 external peer review for acceptance. 



A volume contains about 160 pages, made up by two numbers, published in the Spring and Autumn. 

 Subscriptions may be placed for one or more of the series on an annual basis. Individual numbers 

 and back numbers can be purchased and a Bulletin catalogue, by series, is available. Orders and 

 enquiries should be sent to: 



Intercept Ltd. 

 P.O. Box 716 

 Andover 

 Hampshire SP10 1YG 



Telephone: (0264) 334748 

 Fax: (0264) 334058 



Wo rid List abbreviation: Bull. nat. Hist. Mus. Lond. (Zool.) 

 © The Natural History Museum, 1993 



Zoology Series 

 ISSN 0968 - 0470 Vol. 59, No. 2, pp. 97-170 



The Natural History Museum 



Cromwell Road 



London SW7 5BD Issued 25 November 1993 



Typeset by Ann Buchan (Typesetters), Middlesex 

 Printed in Great Britain at The Alden Press, Oxford 



