108 



Station 3 [no locality]. Brownish mud with lumps 

 of blueish clay throughout, residue about one 

 quarter-pound and floatings small. 

 Station 5 [no locality]. Blue ooze, residue and 

 floatings small. 



Station 9 [no locality]. Results poor. 

 Station 11 [no locality]. 

 Station 12 [no locality]. 



Station 14. Similar to Station 13 [Segaar, New 

 Guinea, coral sand and mud, residue about six 

 ounces, floatings rich]. 

 Station 15 [no locality]. 

 Area 2 Station 17. Muntok Banka, blue mud, residue 

 eight ounces, floatings rich. 



Station 19 [no locality]. Earthy coloured, river- 

 looking mud, few foraminifera. 

 Station 21. Paney, northeast coast of Sumatra. 

 Station 27 [no locality]. 

 Station 28 [no locality]. 



Durrand (1898:257) adds a postscript, stating that ... 'it is 

 important to bear in mind all this series was obtained from 

 shallow water close inshore . . .'. It is clear from the aggluti- 

 nating foraminifera revised here, that most of the localities 

 were in fact brackish, associated with mangroves. 



SYSTEMATIC DESCRIPTIONS 



Order FORAMINIFERIDA Eichwald, 1830 



Suborder TROCHAMMININA Bronnimann & Whittaker, 



1988 



Apart from the hierarchy listed above, no further suprage- 

 neric taxa will be used. Until we can be certain that the 

 families and superfamilies of agglutinating foraminifera used 

 by Loeblich & Tappan (1987) represent homogeneous units 

 with respect to the wall structure, then it is better, for the 

 present, not to use them. Similarly, genera are used in 

 'inverted commas' when the wall structure of their type 

 species has not yet been examined. The eleven species 

 described here, at least, all have a Trochamminina-type wall, 

 defined by Bronnimann & Whittaker (1988) as . . . 'consist- 

 ing of organic and agglutinated phases. Agglutinant bound by 

 organic cement and outer and inner organic sheets. Devoid of 

 perforations or alveolar pseudopores'. 



The synonymies are not meant to be comprehensive, they 

 are selective, merely listing the original reference, junior 

 synonyms, changes of generic combination and important 

 citations from the study area. 



P. BRONNIMANN AND J.E. WHITTAKER 

 Genus ACUPEINA Bronnimann & Zaninetti, 19846 



Type species. Haplophragmium salsum Cushman & Bronni- 

 mann, 1948a (= junior subjective synonym of Haplophrag- 

 mium agglutinans d'Orbigny var. triperforata Millett, 1899). 



Acupeina triperforata (Millett, 1899) Figs 1.2, 13-15 



1899 Haplophragmium agglutinans d'Orbigny var. triper- 

 forata Millett: 358{pars); pi. 5, figs 2a, b (lectotype) 

 only; non figs 3a, b. 



1948a Haplophragmium salsum Cushman & Bronnimann: 

 16,17; pi. 3, figs 10-13. 



1965 Lituola salsa (Cushman & Bronnimann); Bronni- 

 mann & Zaninetti: 608-615; figs 1-3. 



19846 Acupeina salsa (Cushman & Bronnimann); Bronni- 

 mann & Zaninetti: 219-222; figs Al-4, Bl,2. 



19846 Acupeina triperforata (Millett); Bronnimann & 

 Zaninetti: 222 (addendum). 



1988 Acupeina triperforata (Millett); Bronnimann & Whit- 

 taker: 112; pi. 4, figs 1-7. 



Remarks. Millett (1899, pi. 5, figs 2,3; here reproduced as 

 Figs 1.2, 3) illustrated four views of his new variety triperfo- 

 rata. Examination of the original material shows that two 

 different brackish species are involved: Acupeina triperforata 

 (Millett) and Arenoparrella mexicana (Kornfeld). 



The individual drawn by Millett (1899, pi. 5, figs 2a, b; here 

 reproduced in Fig. 1.2a,b) in side and apertural views, has 

 been re-illustrated by SEM in Figs 13-15. The micrographs 

 show side and edge views of a test, initially streptospiral then 

 uniserial, and the detail of the multiple aperture which 

 consists of three closely spaced, virtually equidistant pores (of 

 around 25 um diameter) with upturned rims. Millett appar- 

 ently believed that the aperture of his new variety invariably 

 consisted of the three rounded pores, hence the name. The 

 individual in Figs 13-15 is undoubtedly Millett's figured 

 specimen and is here formally designated lectotype. 



The specimen drawn by Millett (1899, pi. 5, figs 3a, b; 

 reproduced here in Fig, 1.3a,b) in side and apertural views, 

 has been re-illustrated by SEM in Figs 9-12 not only to show 

 both sides of the test but the details of the composite 

 aperture. Its morphology is quite different from the lectotype 

 of H. agglutinans var. triperforata. It represents, in fact, a 

 typical specimen of Arenoparrella mexicana (Kornfeld, 

 1931)(see below). It is unfortunate that Loeblich & Tappan 

 (1987: 21, pi. 71, figs 3,4) illustrated this very specimen, 

 together with the lectotype, as A. triperforata. It is also worth 

 noting that Millett's pi. 5, fig. 3b is the edge view of fig. 3a, 

 but as can be seen from our SEM illustration, rather mislead- 

 ing. It purports to show only three large pores with everted 

 borders. In reality, it has a single oblique-perpendicular slit 

 and 12 small peripheral pores of 5-6 um diameter, devoid of 

 rims. Closer examination of Millett's apertural view (see Fig. 

 1.3b) may just show the termination of the slit (the specimen 

 is tilted forward), but the determination of the pores is still 

 seriously in error. 



Fig. 1.1-1.10, 1.12 Reproduction of part of Plate 5 of Millett (1899). The original identifications were as follows: Fig. 1.1, Haplophragmium 

 agglutinans (d'Orbigny), X112; Fig. 1.2, 3, H. agglutinans var. triperforata var.nov., X112; Fig. 1.4-6, H. cassis (Parker). X112; Fig. 1.7, 

 H. cassis (Parker) or ?Reophax, X75; Fig. 1.8, H. compressum Goes, X75; Fig. 1.9, H. nanum Brady, X112; Fig. 1.10, H. anceps Brady, 

 X56; Fig. 1.12, Trochammina ochracea (Williamson), X75. Reproduced by permission of the Royal Microscopical Society. 



Fig. 2.1 Reproduction of part of Plate 1 of Millett (1900). It was originally identified as Bigenerina digitata d'Orbigny var., X169. 

 Reproduced by permission of the Royal Microscopical Society. 



