112 



P. BRONNIMANN AND J.E. WHITTAKER 



agglutinans, the type species, should have the same wall type, 

 then exiguus would be correctly placed in Ammob acuities. If 

 not, then it would have to be placed in a new genus. In view 

 of these uncertainties, Ammobaculites is placed in inverted 

 commas in our treatment of 'A '. exiguus. 



'Ammobaculites' exiguus Cushman & Bronnimann, 

 19486 Figs 1.1,42-44 



1885 Haplophragmium agglutinans (d'Orbigny); Balkwill 

 & Wright: 330; pi. 13, figs 18?, 19,20 (non d'Orbigny, 

 1846). 



1899 Haplophragmium agglutinans (d'Orbigny); Millett: 

 357, pi. 5, figs la,b. 



1938 Ammobaculites agglutinans (d'Orbigny); Barten- 

 stein:391;fig. 14. 



19486 Ammobaculites exiguus Cushman & Bronnimann: 

 38; pi. 7, figs 7,8. 



19526 Ammobaculites cf. exiguus Cushman & Bronnimann; 

 Parker: 443 ;pl. 1, figs 16,17. 



1952 Ammobaculites agglutinans (d'Orbigny); Rottgardt: 

 180; pi. 1, fig. 4. 1954 Ammobaculites exiguus Cush- 

 man & Bronnimann; Phleger: 633; pi. 1, fig. 5. 



1956 Ammobaculites sp. B, Warren: 139; pi. 1, figs 22-24. 



1957 Ammobaculites exiguus Cushman & Bronnimann; 

 Todd & Bronnimann: 23; pi. 2, fig. 7. 



1962 Ammobaculites exiguus Cushman & Bronnimann; 

 Benda & Puri: 335; pi. 1, fig. 15. 1973 Ammobacu- 

 lites balkwilli Haynes: 25-27; pi. 2, figs 2,3; pi. 29, 

 figs 5,6; text-fig. 4.1-5. 



1978 Ammobaculites dilitatus (sic) Cushman & Bronni- 

 mann; Schafer & Cole: pi. 3, fig. 9 (non Cushman & 

 Bronnimann, 19486). 



1980 Ammobaculites dilatatus Cushman & Bronnimann; 

 Scott & Medioli: 35; pi. 1, figs 9,10. 



71983 Ammobaculites exiguus Cushman & Bronnimann; 

 Haman: 72; pi. 5, figs 1^1. 



1983 Ammobaculites diversus Cushman & Bronnimann; 

 Haman: 72; pi. 4, figs 14,15 (non Cushman & Bron- 

 nimann, 19486). 



1986 Ammobaculites exiguus Cushman & Bronnimann; 

 Bronnimann & Keij: pi. 3, fig. 7. 



Remarks. Millett (1899: pi. 5, figs la,b; here reproduced as 

 Figs l.la,b) illustrated, without description, a typical speci- 

 men of exiguus under the name of Haplophragmium agglutin- 

 ans (d'Orbigny). The same specimen (BMNH no. 

 1955.11.1.1057) is re-illustrated by SEM in our Figs 43,44. 

 The oblique view (Fig. 43) shows the radially-symmetrical 

 areal and terminal aperture, which is larger than in Millett's 

 drawing. It is not bordered by a rim as that shown by 

 Haman's (1983, pi. 5, figs 1-4) 'A. exiguus', which may 

 represent a different species. Millett's specimen has four 

 uniserial chambers which follow from a planispiral, tightly 

 enrolled early test. The agglutinant is coarse and the sutures 

 in the initial portion are not well defined; on the uniserial 

 portion, they are distinct, however, and run perpendicularly 

 to the elongate axis of the test. 



Illustrated in Fig. 42 (BMNH no. 1911.11.1.1058) is a 

 smaller, albeit damaged specimen, which is more typical of 

 the size of the Malay material. Four radial sutures can be 

 recognized in the coiled portion and there are three chambers 

 in the uniserial part; the final chamber is crushed. 



One of us (P.B.) has re-examined the holotype of A. 



exiguus (registration no. 56761) in the U.S. National Museum 

 of Natural History. Its overall morphology corresponds well 

 with Millett's illustrated specimen of H. agglutinans. How- 

 ever, in its uniserial portion there are five chambers and the 

 agglutinant is finer than in the Malay specimen. Nevertheless, 

 the two both have a circular transverse section and a large 

 radially-symmetrical, terminal aperture without a rim; the 

 intercameral sutures run perpendicular to the elongate axis of 

 the test, there being no suggestion of Ammotium-type 

 sutures. In addition to the holotype of exiguus, there are two 

 slides with paratypes: in slide no. 56762 there is a single 

 paratype; under no. 56763 there are, amongst typical speci- 

 mens, some very small individuals which differ from the type 

 by their thin, elongate tests. These latter have also been 

 encountered by us in the mangrove sediments of Acupe, 

 Brazil. They represent a new species of brackish 'Ammobacu- 

 lites' which will be published elsewhere. It should be noted 

 that 'A', exiguus and this new, minute species, are the only 

 true representatives of 'Ammobaculites' occurring in brackish 

 waters. 



Dimensions of figured specimen (BMNH no. 

 1911.11.1.1057). Height of test — 385 \im; diameter of initial 

 planispiral portion — 135 um; diameter of final chamber — 

 125 ^m; diameter of aperture — 50 \im. 



Environment. In the Millett Collection, specimens are 

 labelled 'Haplophragmium agglutinans' from stations 

 2,9,12,14,15,19,21 and 27; Millett notes (p. 358) that . . . 'the 

 specimens are all minute, and although they occur at most of 

 the Stations, are not very numerous'. According to Parker et 

 al. (1953), 'A', exiguus is a species which lives in brackish as 

 well as in marine waters. 



Genus AMMOTIUM Loeblich & Tappan, 1953 



TYPE SPECIES. Lituola cassis Parker (in Dawson), 1870. 



REMARKS. Ammomarginulina Wiesner, 1931 (type species: 

 A. ensis Wiesner, 1931) is a deep-water genus, with a 

 morphology close to that of the supposedly exclusively 

 brackish-water genus, Ammotium. After having compared 

 the definitions of Ammomarginulina and of Ammotium in 

 Loeblich & Tappan (1987), the question arises as to whether 

 the two are really synonymous. The sutures of the former are, 

 however, less slanting that those of Ammotium, and the test 

 is strongly compressed. Of the shape of the aperture of 

 Ammomarginulina ensis nothing is known except for the fact 

 that it is rounded. Small morphological differences such as 

 these may not be considered sufficient to retain the two 

 genera. However, they seem to represent two disparate 

 homogeneous environmental groups which, should this be 

 sustained, must be separated taxonomically, even if the 

 morphological differences were even less pronounced (see 

 also Resig's (1982: 977-978, pi. 1, figs 3-5,9) description of 

 Ammomarginulina hadalensis Resig from the Peru-Chile 

 Trench, depth 5846 m). Clearly, the wall structure of Ammo- 

 marginulina must also be investigated. 



Ammotium morenoi (Acosta, 1940) 

 1899 



Figs 32-34, 54 



Haplophragmium cassis (Parker): 359 (pars) (non 

 Lituola cassis Parker, 1870). 

 1940 Ammobaculites morenoi Acosta: 272; pi. 49, figs 3,8 

 (holotype) only (non Fig. 1). 



