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J.D. TAYLOR, Y.I. KANTOR AND A.V. SYSOEV 



Ancestor 

 C lav us U 

 Splendnllia C 



Hastula B 

 Pervicacia T 

 Ouphcana C 

 Pseudomelatoma 

 Toxichonella T • 

 Strict is pir a P — 

 Clavatula D 

 Clionella C 

 Clavulata S 

 Turncula N 

 Funa L 



Vexitomina G 

 Pilsbryspira N 

 Aforia A 



Drilliidae 

 Terebridae 



Pseudomelatomidae 



Clavatulinae 



Stnctispindae 



Clavatulinae 



Lophiotoma L 

 Polystira A 

 Micantapex P 

 Borsonia 

 Ophiodermella _ 

 Oenopota L — 

 Tropidoturns F 

 Ananthma M 

 Glyphostoma C — 

 Genota N — i 

 Benthofascis _J 

 Conus V 



Tomopleura V — 

 Eucithara S 

 Mangelia N 

 Mangelia P — 

 Thatcheria M 

 Philbertia P 

 Philbertia L 

 Daphnella R 

 Gymnobela E 

 Teretiopsis L 

 Abyssobela A 

 Taranis M — 



Crassispirmae 



Zonulispirmae 

 — Cochlespinnae 



Turnnae 



Clathurelhnae 



• Oenopotinae 

 Clathurelhnae 



Conorbinae 



Coninae 



Clathurelhnae 



Mangehinae 



Daphnelhnae 



Taraninae 



Fig. 27 Majority-rule (50%) consensus tree. Autapomorphies for each node given in Table 4. Higher taxa names at the top of branches 

 reflect our new classification. Taxon abbreviations in order top to bottom on the tree: Clavus U = Clavus unizonalis, Splendrillia C = 

 Spendrillia chathamensis , Hastula B = Hastula bacillus, Duplicaria C = Duplicaria colorata, Pseudomelatoma P = Pseudomelatoma 

 penicillatus, Toxichonella T = Toxichonella tumida, Strictispira P = Strictispira paxillus , Clavatula D = Clavatula diadema, Clionella S = 

 Clionella sinuata, Clavatula C = Clavatula caerulea, Turricula N = Turricula nelliae, Funa L = Funa latisinuata, Vexitomina G = 

 Vexitomina garrardi, Pilsbryspira N = Pilsbryspira nympha, Aforia A = Aforia abyssalis, Lophiotoma L = Lophiotoma leucotropis , 

 Polystira A = Polystira albida, Micantapex P = Micantapex parengonius, Borsonia O = Borsonia ochraea, Ophiodermella I = 

 Ophiodermella inermis, Oenopota L = Oenopota levidensis, Tropidoturris F = Tropidoturris fossata, Anarithma M = Anarithma metula, 

 Glyphostoma C = Glyphostoma Candida, Genota N = Genota nicklesi, Benthofascis = Benthofascis biconica, Conus V = Conus 

 ventricosus, Tomopleura V = Tomopleura reevei, Eucithara S = Eucithara stromboides, Mangelia N = Mangelia nebula, Mangelia P = 

 Mangelia powisiana, Thatcheria M = Thatcheria mirabilis, Philbertia P = Philbertia purpurea, Philbertia L = Philbertia linearis, Daphnella 

 R = Daphnella reeveana, Gymnobela E = Gymnobela emertoni, Teretiopsis L = Teretiopsis levicarinatus, Abyssobela A = Abyssobela 

 atoxica, Taranis M = Taranis moerchi. 



ribbon and loss of the odontophore. 



Clade 19 is made up of various taxa formerly included in 

 the Borsoniinae and Clathurelhnae with the addition of 

 Oenopota (Oenopotinae). The apomorphies defining the 

 nodes are very unsatisfactory with many reversals. More 

 characters need to be analysed in these taxa to achieve better 

 resolution. 



Borsonia and Ophiodermella (Node 21) have posteriorly 

 situated rhynchodeal sphincters, and fusiform shells. The 



taxa in the other clade (Node 22) have extensible buccal lips 

 and hollow radular teeth with large bases. Although the 

 Oenopotinae have been previously thought to have close 

 affinities with the Mangehinae, they do have acinous salivary 

 glands, rather than the tubular type associated with the latter 

 subfamily. 



A clade comprising Anarithma and Glyphostoma is defined 

 (Node 24) by three characters; a posteriorly situated rhyn- 

 chodeal sphincter, a single salivary duct and apertural orna- 



